#NEXUS BEGIN TAXA; Dimensions NTax = 283; TaxLabels 'Phrynomantis microps' 'Hemisus sudanensis' 'Hemisus marmoratus' 'Balebreviceps hillmani' 'Breviceps mossambicus' 'Breviceps adspersus' 'Breviceps montanus' 'Breviceps fuscus' 'Breviceps gibbosus' 'Breviceps macrops' 'Breviceps namaquensis' 'Breviceps branchi' 'Spelaeophryne methneri' 'Probreviceps loveridgei' 'Probreviceps uluguruensis' 'Probreviceps durirostris' 'Probreviceps sp Nguru' 'Probreviceps sp Rubeho' 'Probreviceps sp Kigogo' 'Probreviceps sp Udzungwa' 'Probreviceps rungwensis' 'Probreviceps macrodactylus' 'Callulina shengena' 'Callulina laphami' 'Callulina dawida' 'Callulina kanga' 'Callulina sp lowland' 'Callulina sp Rubeho' 'Callulina hanseni' 'Callulina meteora' 'Callulina stanleyi' 'Callulina kisiwamsitu' 'Callulina kreffti' 'Acanthixalus spinosus' 'Acanthixalus sonjae' 'Semnodactylus wealii' 'Paracassina obscura' 'Paracassina kounhiensis' 'Phlyctimantis maculata' 'Phlyctimantis keithae' 'Phlyctimantis verrucosus' 'Phlyctimantis boulengeri' 'Phlyctimantis leonardi' 'Kassina kuvangensis' 'Kassina fusca' 'Kassina lamottei' 'Kassina schioetzi' 'Kassina cassinoides' 'Kassina senegalensis' 'Kassina decorata' 'Kassina maculosa' 'Kassina arboricola' 'Kassina cochranae' 'Opisthothylax immaculatus' 'Afrixalus enseticola' 'Tachycnemis seychellensis' 'Heterixalus madagascariensis' 'Heterixalus boettgeri' 'Heterixalus alboguttatus' 'Heterixalus punctatus' 'Heterixalus andrakata' 'Heterixalus tricolor' 'Heterixalus variabilis' 'Heterixalus luteostriatus' 'Heterixalus betsileo' 'Heterixalus carbonei' 'Heterixalus rutenbergi' 'Afrixalus vibekensis' 'Afrixalus weidholzi' 'Afrixalus lacustris' 'Afrixalus dorsimaculatus 1' 'Afrixalus morerei' 'Afrixalus dorsimaculatus 2' 'Afrixalus knysae' 'Afrixalus spinifrons' 'Afrixalus delicatus' 'Afrixalus brachycnemis' 'Afrixalus stuhlmanni' 'Afrixalus sylvaticus' 'Afrixalus laevis' 'Afrixalus lacteus' 'Afrixalus fornasini' 'Afrixalus equatorialis' 'Afrixalus wittei' 'Afrixalus osorioi' 'Afrixalus paradorsalis 3' 'Afrixalus paradorsalis 2' 'Afrixalus paradorsalis 1' 'Afrixalus nigeriensis' 'Afrixalus vittiger 1' 'Afrixalus vittiger 2' 'Afrixalus dorsalis 3' 'Afrixalus dorsalis 1' 'Afrixalus quadrivittatus 1' 'Afrixalus quadrivittatus 2' 'Afrixalus dorsalis 2' 'Afrixalus fulvovittatus 2' 'Afrixalus fulvovittatus 1' 'Morerella cyanophthalma' 'Cryptothylax greshoffii' 'Hyperolius semidiscus' 'Hyperolius lupiroensis' 'Hyperolius parkeri' 'Hyperolius viridis' 'Hyperolius pusillus' 'Hyperolius benguellensis' 'Hyperolius inyangae' 'Hyperolius nasutus' 'Hyperolius dartevellei' 'Hyperolius rwandae' 'Hyperolius friedemanni' 'Hyperolius howelli' 'Hyperolius acuticeps' 'Hyperolius poweri' 'Hyperolius igbettensis' 'Hyperolius jacobseni' 'Hyperolius adspersus' 'Hyperolius pseudoargus' 'Hyperolius fusciventris 2' 'Hyperolius fusciventris 1' 'Hyperolius bolifambae' 'Hyperolius ocellatus 1' 'Hyperolius ocellatus 2' 'Hyperolius sp nov 1' 'Hyperolius camerunensis' 'Hyperolius riggenbachi' 'Hyperolius guttulatus 2' 'Hyperolius guttulatus 1' 'Hyperolius phantasticus' 'Hyperolius pardalis' 'Hyperolius steindachneri' 'Hyperolius argus' 'Hyperolius bocagei' 'Hyperolius lamottei' 'Hyperolius tuberculatus' 'Hyperolius dintelmanni' 'Hyperolius parallelus parallelus' 'Hyperolius parallelus marginatus' 'Hyperolius parallelus alborufus' 'Hyperolius parallelus pyrrhodictyon' 'Hyperolius marmoratus taeniatus' 'Hyperolius marmoratus marmoratus' 'Hyperolius marmoratus swynnertoni' 'Hyperolius mariae' 'Hyperolius reesi' 'Hyperolius nitidulus 1' 'Hyperolius spatzi' 'Hyperolius nitidulus 2' 'Hyperolius viridiflavus pitmani' 'Hyperolius viridiflavus sp' 'Hyperolius viridiflavus destefanii' 'Hyperolius viridiflavus variablilis' 'Hyperolius viridiflavus ngorongoriensis' 'Hyperolius viridiflavus ommatostictus' 'Hyperolius viridiflavus pantherinus' 'Hyperolius viridiflavus glandicolor' 'Chlorolius koehleri' 'Hyperolius tuberilinguis' 'Hyperolius mosaicus' 'Hyperolius endjami 2' 'Hyperolius endjami 1' 'Hyperolius cystocandicans' 'Hyperolius jackie' 'Hyperolius discodactylus' 'Hyperolius lateralis' 'Hyperolius castaneus' 'Hyperolius frontalis' 'Hyperolius platyceps' 'Hyperolius cinereus' 'Hyperolius raymondii' 'Hyperolius langi' 'Hyperolius kuligae' 'Hyperolius ademetzi' 'Hyperolius tanneri' 'Hyperolius minitissimus' 'Hyperolius burgessi' 'Hyperolius spinigularis' 'Hyperolius torrentis' 'Hyperolius chlorosteus' 'Hyperolius occidentalis' 'Hyperolius sylvaticus' 'Hyperolius baumanni' 'Hyperolius picturatus' 'Hyperolius zonatus' 'Hyperolius bobirensis' 'Hyperolius concolor 2' 'Hyperolius concolor 1' 'Hyperolius concolor 3' 'Hyperolius montanus' 'Hyperolius pictus' 'Hyperolius substriatus' 'Hyperolius sp nov 2' 'Hyperolius mitchelli' 'Hyperolius rubrovermiculatus' 'Hyperolius quinquevittatus' 'Hyperolius kivuensis 1' 'Hyperolius kivuensis 3' 'Hyperolius kivuensis 2' 'Hyperolius balfouri 1' 'Hyperolius balfouri 2' 'Hyperolius veithi' 'Hyperolius cinnamomeoventris 2' 'Hyperolius cinnamomeoventris 1' 'Hyperolius cinnamomeoventris 3' 'Hyperolius thomensis' 'Hyperolius molleri' 'Nyctibates corrugatus' 'Scotobleps gabonicus' 'Astylosternus laticephalus' 'Astylosternus occidentalis' 'Trichobatrachus robustus' 'Astylosternus diadematus' 'Astylosternus schioetzi' 'Astylosternus batesi' 'Leptopelis parkeri' 'Leptopelis macrotis' 'Leptopelis millsoni' 'Leptopelis rufus' 'Leptopelis argenteus' 'Leptopelis yaldeni' 'Leptopelis vannutellii' 'Leptopelis susanae' 'Leptopelis gramineus' 'Leptopelis kivuensis' 'Leptopelis ocellatus' 'Leptopelis spiritusnoctis' 'Leptopelis viridis' 'Leptopelis aubryi' 'Leptopelis natalensis' 'Leptopelis palmatus' 'Leptopelis calcaratus' 'Leptopelis brevirostris' 'Leptopelis notatus' 'Leptopelis modestus' 'Leptopelis flavomaculatus' 'Leptopelis nordequatorialis' 'Leptopelis bocagii' 'Leptopelis vermiculatus' 'Leptopelis aubryioides' 'Leptopelis boulengeri' 'Leptopelis fiziensis' 'Leptopelis karissimbensis' 'Cardioglossa melanogaster' 'Cardioglossa gracilis' 'Cardioglossa elegans' 'Cardioglossa leucomystax' 'Cardioglossa occidentalis' 'Cardioglossa pulchra' 'Cardioglossa oreas' 'Cardioglossa manengouba' 'Cardioglossa escalerae' 'Cardioglossa trifasciata' 'Cardioglossa annulata' 'Cardioglossa nigromaculata' 'Arthroleptis schubotzi' 'Arthroleptis xenodactyloides' 'Arthroleptis fichika' 'Arthroleptis sylvaticus' 'Arthroleptis taeniatus' 'Arthroleptis reichei' 'Arthroleptis affinis' 'Arthroleptis nikeae' 'Arthroleptis tanneri' 'Arthroleptis francei' 'Arthroleptis stenodactylus' 'Arthroleptis perreti' 'Arthroleptis palava' 'Arthroleptis variabilis' 'Arthroleptis adelphus' 'Arthroleptis brevipes' 'Arthroleptis bioko' 'Arthroleptis poecilonotus' 'Leptodactylodon mertensi' 'Leptodactylodon erythrogaster' 'Leptodactylodon perreti' 'Leptodactylodon axillaris' 'Leptodactylodon polyacanthus' 'Leptodactylodon bicolor' 'Leptodactylodon bueanus' 'Leptodactylodon boulengeri' 'Leptodactylodon ventrimarmoratus' 'Leptodactylodon ovatus' 'Leptodactylodon ornatus' ; END; BEGIN TREES; Tree tree1 = ('Phrynomantis microps':0.178054,(((('Hemisus sudanensis':0.011442,'Hemisus marmoratus':0.003714):0.252643,('Balebreviceps hillmani':0.144157,((('Breviceps mossambicus':0.031189,'Breviceps adspersus':0.057508):0.07571,('Breviceps montanus':0.029545,('Breviceps fuscus':0.021642,('Breviceps gibbosus':0.034182,('Breviceps macrops':0.028471,('Breviceps namaquensis':0.030869,'Breviceps branchi':0.019592):0.013673):0.031059):0.002364):0.006903):0.021027):0.053239,(('Spelaeophryne methneri':0.112935,(('Probreviceps loveridgei':0.048756,'Probreviceps uluguruensis':0.019845):0.042663,(('Probreviceps durirostris':0.00761,'Probreviceps sp Nguru':0.015379):0.016956,('Probreviceps sp Rubeho':0.026481,('Probreviceps sp Kigogo':0.008002,('Probreviceps sp Udzungwa':0.005902,('Probreviceps rungwensis':1e-06,'Probreviceps macrodactylus':0.003569):0.001754):0.003753):0.01421):1e-06):0.010539):0.051728):0.013571,(('Callulina shengena':0.006376,'Callulina laphami':0.05275):0.017848,('Callulina dawida':0.021404,('Callulina kanga':0.01762,('Callulina sp lowland':0.01536,('Callulina sp Rubeho':0.03052,(('Callulina hanseni':0.011747,'Callulina meteora':0.012663):0.011102,('Callulina stanleyi':0.031681,('Callulina kisiwamsitu':0.005336,'Callulina kreffti':0.003086):0.014516):0.027264):1e-06):0.004363):1e-06):0.009483):0.016251):0.047511):0.064801):0.015308):0.050037):0.070339,(((('Acanthixalus spinosus':0.003682,'Acanthixalus sonjae':0.009857):0.152811,('Semnodactylus wealii':0.084505,(('Paracassina obscura':0.089491,'Paracassina kounhiensis':0.028892):0.032634,(('Phlyctimantis maculata':0.041077,(('Phlyctimantis keithae':0.033491,'Phlyctimantis verrucosus':0.022033):0.009613,('Phlyctimantis boulengeri':0.019258,'Phlyctimantis leonardi':0.021238):0.010937):0.003021):0.036966,('Kassina kuvangensis':0.056783,(('Kassina fusca':0.043211,('Kassina lamottei':0.03216,('Kassina schioetzi':0.011999,'Kassina cassinoides':0.030985):0.012476):0.002108):0.01981,('Kassina senegalensis':0.036342,(('Kassina decorata':0.005271,'Kassina maculosa':0.009684):0.006176,('Kassina arboricola':0.009479,'Kassina cochranae':0.009891):0.003215):0.026396):0.011858):0.012894):0.019375):0.015102):0.00359):0.071695):0.009358,('Opisthothylax immaculatus':0.124173,((('Afrixalus enseticola':0.067375,('Tachycnemis seychellensis':0.043145,(('Heterixalus madagascariensis':0.019025,('Heterixalus boettgeri':0.010733,'Heterixalus alboguttatus':0.017205):0.004096):0.012662,('Heterixalus punctatus':0.030275,(('Heterixalus andrakata':0.005655,('Heterixalus tricolor':0.028936,'Heterixalus variabilis':1e-06):0.006187):0.025082,('Heterixalus luteostriatus':0.066752,('Heterixalus betsileo':0.013446,('Heterixalus carbonei':0.018203,'Heterixalus rutenbergi':0.013627):0.003721):1e-06):0.012761):0.004927):0.008055):0.0141):0.033091):0.002244,((('Afrixalus vibekensis':0.072972,('Afrixalus weidholzi':0.041999,('Afrixalus lacustris':0.036175,('Afrixalus dorsimaculatus 1':0.037718,('Afrixalus morerei':0.023852,'Afrixalus dorsimaculatus 2':0.015307):0.007058):0.006742):1e-06):0.005711):0.003308,(('Afrixalus knysae':0.012612,'Afrixalus spinifrons':0.01132):0.025783,('Afrixalus delicatus':0.038702,('Afrixalus brachycnemis':0.024595,('Afrixalus stuhlmanni':0.01029,'Afrixalus sylvaticus':0.016901):0.010525):0.001921):0.007953):0.009852):0.013379,(('Afrixalus laevis':0.029134,'Afrixalus lacteus':0.033516):0.019809,('Afrixalus fornasini':0.049644,(('Afrixalus equatorialis':0.020501,('Afrixalus wittei':0.007301,'Afrixalus osorioi':0.016069):0.0057):0.023965,(('Afrixalus paradorsalis 3':0.021251,('Afrixalus paradorsalis 2':0.03881,'Afrixalus paradorsalis 1':0.021277):0.00782):0.022532,(('Afrixalus nigeriensis':0.033986,('Afrixalus vittiger 1':0.014681,'Afrixalus vittiger 2':0.013814):0.015756):0.012296,(('Afrixalus dorsalis 3':0.035815,'Afrixalus dorsalis 1':0.022078):0.005159,(('Afrixalus quadrivittatus 1':0.03403,'Afrixalus quadrivittatus 2':0.03936):0.018511,('Afrixalus dorsalis 2':0.031843,('Afrixalus fulvovittatus 2':0.025256,'Afrixalus fulvovittatus 1':0.036296):0.005705):0.010377):0.004988):0.004279):0.024392):0.001315):0.014543):0.005847):0.017429):0.016519):0.028387,(('Morerella cyanophthalma':0.053853,'Cryptothylax greshoffii':0.089737):0.028044,('Hyperolius semidiscus':0.098559,(('Hyperolius lupiroensis':0.01233,'Hyperolius parkeri':1e-06):0.074866,((('Hyperolius viridis':0.039901,'Hyperolius pusillus':0.037961):0.007329,(('Hyperolius benguellensis':0.03004,'Hyperolius inyangae':0.025641):0.009198,('Hyperolius nasutus':0.005763,('Hyperolius dartevellei':0.015376,('Hyperolius rwandae':0.01318,('Hyperolius friedemanni':0.008092,(('Hyperolius howelli':0.002181,'Hyperolius acuticeps':0.001952):0.018981,(('Hyperolius poweri':0.013424,'Hyperolius igbettensis':0.041208):0.003962,('Hyperolius jacobseni':0.006762,'Hyperolius adspersus':0.006246):0.010055):0.001538):1e-06):0.003005):0.001665):1e-06):0.006928):0.015974):0.046951,((('Hyperolius pseudoargus':0.048775,(('Hyperolius fusciventris 2':0.012415,'Hyperolius fusciventris 1':0.022241):0.032541,('Hyperolius bolifambae':0.037674,(('Hyperolius ocellatus 1':0.016148,'Hyperolius ocellatus 2':0.021607):0.01576,('Hyperolius sp nov 1':0.03178,('Hyperolius camerunensis':0.023948,'Hyperolius riggenbachi':0.023329):0.012593):0.001855):0.00543):0.017172):0.001543):0.005233,(('Hyperolius guttulatus 2':0.016718,'Hyperolius guttulatus 1':0.019583):0.034505,(('Hyperolius phantasticus':0.014384,('Hyperolius pardalis':0.016383,'Hyperolius steindachneri':0.014717):0.005933):0.020056,('Hyperolius argus':0.043432,('Hyperolius bocagei':0.047638,('Hyperolius lamottei':0.034607,((('Hyperolius tuberculatus':0.028279,'Hyperolius dintelmanni':0.012171):0.005097,('Hyperolius parallelus parallelus':0.020796,('Hyperolius parallelus marginatus':0.011746,('Hyperolius parallelus alborufus':0.02375,'Hyperolius parallelus pyrrhodictyon':0.017829):0.004458):0.007454):0.001942):0.006244,(('Hyperolius marmoratus taeniatus':0.016578,(('Hyperolius marmoratus marmoratus':0.007464,'Hyperolius marmoratus swynnertoni':0.008809):0.009346,('Hyperolius mariae':0.014967,'Hyperolius reesi':0.021829):0.016406):0.000555):0.006568,(('Hyperolius nitidulus 1':0.021782,('Hyperolius spatzi':0.027716,'Hyperolius nitidulus 2':0.014022):0.005625):0.021482,(('Hyperolius viridiflavus pitmani':0.015268,('Hyperolius viridiflavus sp':0.022437,('Hyperolius viridiflavus destefanii':0.017503,'Hyperolius viridiflavus variablilis':0.024752):0.015763):0.001874):0.004393,('Hyperolius viridiflavus ngorongoriensis':0.009245,('Hyperolius viridiflavus ommatostictus':0.011928,('Hyperolius viridiflavus pantherinus':0.01609,'Hyperolius viridiflavus glandicolor':0.011076):0.003756):0.006271):0.001714):0.004221):0.001439):0.000934):0.010043):0.014778):0.0032):0.001272):0.004243):0.010355):0.032008,('Chlorolius koehleri':0.043767,('Hyperolius tuberilinguis':0.068262,((('Hyperolius mosaicus':0.008695,('Hyperolius endjami 2':0.012027,'Hyperolius endjami 1':0.007244):0.007534):0.028688,('Hyperolius cystocandicans':0.035586,('Hyperolius jackie':0.024578,(('Hyperolius discodactylus':0.022702,'Hyperolius lateralis':0.012335):0.010667,('Hyperolius castaneus':0.017932,'Hyperolius frontalis':0.023167):0.003725):0.009553):0.003186):0.007411):0.011334,((('Hyperolius platyceps':0.046779,('Hyperolius cinereus':0.029669,'Hyperolius raymondii':0.025563):0.014779):0.023447,('Hyperolius langi':0.050024,('Hyperolius kuligae':0.021661,'Hyperolius ademetzi':0.030238):0.012435):0.018803):1e-06,(('Hyperolius tanneri':0.021694,('Hyperolius minitissimus':0.024461,('Hyperolius burgessi':0.014326,'Hyperolius spinigularis':0.022374):0.005613):0.002209):0.038062,(('Hyperolius torrentis':0.03497,(('Hyperolius chlorosteus':0.009946,'Hyperolius occidentalis':0.009987):0.015823,('Hyperolius sylvaticus':0.012301,('Hyperolius baumanni':0.012727,'Hyperolius picturatus':0.008687):0.012955):0.012678):0.016589):0.007509,(((('Hyperolius zonatus':0.024733,'Hyperolius bobirensis':0.013783):0.003989,('Hyperolius concolor 2':0.019653,('Hyperolius concolor 1':0.019977,'Hyperolius concolor 3':0.013833):0.013407):0.004129):0.019931,('Hyperolius montanus':0.019801,(('Hyperolius pictus':0.029378,'Hyperolius substriatus':0.02819):0.005333,('Hyperolius sp nov 2':0.019428,('Hyperolius mitchelli':0.019941,'Hyperolius rubrovermiculatus':0.010944):0.005538):0.021072):0.002873):0.023255):0.002441,(('Hyperolius quinquevittatus':0.04128,(('Hyperolius kivuensis 1':0.019397,'Hyperolius kivuensis 3':0.031653):0.016117,('Hyperolius kivuensis 2':0.030809,('Hyperolius balfouri 1':0.009216,'Hyperolius balfouri 2':0.01621):0.027206):0.010039):0.007023):0.013252,(('Hyperolius veithi':0.019139,('Hyperolius cinnamomeoventris 2':0.012984,'Hyperolius cinnamomeoventris 1':0.007668):0.014087):0.010112,('Hyperolius cinnamomeoventris 3':0.03108,('Hyperolius thomensis':0.007123,'Hyperolius molleri':0.008506):0.012027):0.007823):0.027795):0.003126):0.007999):0.012285):0.005176):0.011907):0.004396):0.00599):0.035257):0.012486):0.003331):0.021949):0.020205):0.003485):0.018968):0.05364):0.129092,(('Nyctibates corrugatus':0.111402,('Scotobleps gabonicus':0.144019,(('Astylosternus laticephalus':0.025024,'Astylosternus occidentalis':0.008697):0.065501,('Trichobatrachus robustus':0.044798,('Astylosternus diadematus':0.035575,('Astylosternus schioetzi':0.004295,'Astylosternus batesi':0.010784):0.004685):0.03898):0.024187):0.026018):0.013073):0.015208,((('Leptopelis parkeri':0.134229,(('Leptopelis macrotis':0.015404,'Leptopelis millsoni':0.009442):0.011613,('Leptopelis rufus':0.030196,(('Leptopelis argenteus':0.0416,('Leptopelis yaldeni':0.008719,('Leptopelis vannutellii':0.004486,('Leptopelis susanae':0.022557,'Leptopelis gramineus':0.015577):0.010439):1e-06):0.034591):0.00534,(('Leptopelis kivuensis':0.029392,('Leptopelis ocellatus':0.033451,('Leptopelis spiritusnoctis':0.057045,('Leptopelis viridis':0.039779,'Leptopelis aubryi':0.019707):0.006999):0.032535):0.013824):0.005611,('Leptopelis natalensis':0.038264,(('Leptopelis palmatus':0.023647,('Leptopelis calcaratus':0.038983,('Leptopelis brevirostris':0.027309,'Leptopelis notatus':0.030858):0.00915):0.003314):0.004391,(('Leptopelis modestus':0.032613,('Leptopelis flavomaculatus':0.022344,('Leptopelis nordequatorialis':0.011486,'Leptopelis bocagii':0.008476):0.005659):0.005313):0.004465,(('Leptopelis vermiculatus':0.031151,'Leptopelis aubryioides':0.052456):0.00993,('Leptopelis boulengeri':0.036326,('Leptopelis fiziensis':0.011153,'Leptopelis 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