((('Moraria insignipes',('Bryocamptus nivalis','Bryocamptus vejdovskyi')),('Bryocamptus arcticus','Moraria duthei')),'Canthocamptus staphylinus');
((('Attheyella crassa','Attheyella dentata'),('Attheyella nordenskioldii I','Attheyella nordenskioldii II')),'Canthocaptus staphylinus');
((('Moraria mrazeki','Moraria brevipes'),'Moraria duthei'),'Canthocamptus staphylinus');