((('Mucropedia cookorum',('Zeusodes septimus',(('Zeusodes arenicolus 5','Neozausodes shulenbergeni'),(('Zeusodes arenicolus 2','Zeusodes arenicolus 1'),('Zeusodes arenicolus 4','Zeusodes arenicolus 3')))))),'Tigriopus japonicus','Tigriopus californicus');