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(2019). Revealing higher than expected diversity of Harpacticoida (Crustacea:Copepoda) in the North Sea using MALDI-TOF MS and molecular barcoding. Scientific Reports, 9(1). https://doi.org/10.1038/s41598-019-45718-7 2019 Cletodes longicaudata SR 641 2949782 Stylicletodes longicaudatus Cletodes longicaudata SR 430 Longipedia coronata SR 189 2954944 Longipedia coronata Longipedia coronata DP 085 Longipedia coronata SR 242 Longipedia coronata SR 241 Longipedia coronata SR 180 Longipedia coronata SR 243 Longipedia coronata DP 084 Longipedia coronata DP 086 Longipedia coronata DP 123 Nannopus palustris DP 007 1020747 Nannopus palustris Cletodes longicaudata DP 263 Nannopus palustris DP 057 Nannopus palustris DP 083 Nannopus palustris SR 449 Nannopus palustris DP 022 Nannopus palustris SR 451 Nannopus palustris SR 452 Nannopus palustris SR 476 Nannopus palustris DP 021 Nannopus palustris SR 456 Nannopus palustris SR 516 Cletodes longicaudata DP 271 Nannopus palustris SR 478 Spinapecruris curvirostris SR 606 2949736 Spinapecruris curvirostris Spinapecruris curvirostris DP 092 Spinapecruris curvirostris DP 125 Spinapecruris curvirostris DP 091 Spinapecruris curvirostris SR 422 Spinapecruris curvirostris SR 706 Spinapecruris curvirostris SR 687 Spinapecruris curvirostris SR 713 Spinapecruris curvirostris DP 090 Cletodes longicaudata DP 270 Platychelipus littoralis DP 023 2950244 Platychelipus littoralis Platychelipus littoralis SR 455 Platychelipus littoralis DP 019 Platychelipus littoralis SR 453 Platychelipus littoralis SR 462 Platychelipus littoralis DP 047 Platychelipus littoralis SR 460 Asellopsis hispida SR 587 2950080 Asellopsis hispida Asellopsis intermedia DP 048 2950090 Asellopsis intermedia Asellopsis intermedia SR 261 Cletodes limicola SR 132 2949765 Cletodes limicola Asellopsis intermedia DP 001 Asellopsis intermedia SR 274 Asellopsis intermedia SR 275 Laophonte longicaudata SR 036 2950143 Laophonte longicaudata Laophonte longicaudata SR 148 Laophonte longicaudata SR 236 Laophonte longicaudata SR 149 Laophonte longicaudata DP 260 Laophonte longicaudata DP 261 Laophonte longicaudata SR 237 Cletodes limicola SR 133 2949765 Cletodes limicola Heteropsyllus curticaudatus SR 178 2953532 Heteropsyllus curticaudatus Heteropsyllus curticaudatus SR 646 Heteropsyllus curticaudatus DP 320 Heteropsyllus curticaudatus DP 322 Heteropsyllus curticaudatus DP 268 Heteropsyllus curticaudatus DP 104 Heteropsyllus curticaudatus DP 319 Heteropsyllus curticaudatus DP 321 Heteropsyllus curticaudatus DP 105 Heteropsyllus curticaudatus SR 144 Cletodes limicola SR 225 Heteropsyllus curticaudatus SR 117 Heteropsyllus curticaudatus SR 119 Heteropsyllus curticaudatus SR 143 Aquilastacus n sp 1 SR 287 2954596 Aquilastacus Aquilastacus n sp 2 SR 288 Paraleptastacus espinulatus 2 DP 115 /transl table 5 /transl table 5 2954043 Paraleptastacus espinulatus Paraleptastacus espinulatus DP 117 Paraleptastacus espinulatus SR 417 Paraleptastacus espinulatus SR 255 Paraleptastacus espinulatus DP 162 Cletodes limicola SR 110 Paraleptastacus espinulatus SR 423 Paraleptastacus espinulatus SR 574 Paraleptastacus espinulatus SR 424 Paraleptastacus espinulatus DP 167 Paraleptastacus espinulatus SR 600 Paraleptastacus espinulatus DP 120 Paraleptastacus espinulatus SR 671 Paraleptastacus espinulatus SR 256 Paraleptastacus espinulatus DP 114 Paraleptastacus espinulatus DP 161 Cletodes limicola SR 408 Paraleptastacus espinulatus DP 119 Paraleptastacus espinulatus SR 659 Cletodes tenuipes DP 106 2949743 Cletodes tenuipes Cletodes tenuipes DP 108 Cletodes tenuipes SR 616 Cletodes tenuipes SR 116 Cletodes tenuipes SR 111 Cletodes n sp SR 128 2949637 Cletodes Cletodes n sp SR 710 Cletodes n sp DP 298 Cletodes limicola DP 292 Cletodes n sp SR 711 Leptastacus macronyx complex 2 SR 586 2954473 Leptastacus macronyx Leptastacus macronyx complex 2 SR 571 Leptastacus macronyx complex 2 SR 657 Leptastacus macronyx complex 2 SR 658 Leptastacus macronyx complex 1 SR 656 Leptastacus cf laticaudatus 2 SR 679 2954939 Leptastacus laticaudatus Leptastacus cf laticaudatus 2 SR 196 Leptastacus cf laticaudatus 2 SR 197 Leptastacus cf laticaudatus 3 DP 121 Cletodes longicaudata SR 642 Cletodes limicola DP 269 Leptastacus cf laticaudatus 3 SR 577 Leptastacus cf laticaudatus 3 DP 122 Leptastacus cf laticaudatus 3 SR 570 Leptastacus cf laticaudatus 3 DP 118 Leptastacus cf laticaudatus 1 SR 569 Leptastacus cf laticaudatus 1 SR 182 Leptastacus cf laticaudatus 1 SR 591 Leptastacus cf laticaudatus 1 DP 195 Leptastacus cf laticaudatus 1 SR 573 Leptastacus cf laticaudatus 1 DP 198 Cletodes limicola DP 262 Leptastacus cf laticaudatus 1 SR 581 Leptastacidae cf gen Minervella SR 672 Rhizothrix curvata DP 137 2954610 Rhizothrix curvata Rhizothrix curvata SR 168 Rhizothrix curvata SR 167 Rhizothrix curvata DP 136 Rhizothrix curvata SR 248 Rhizothrix curvata SR 166 Rhizothrix curvata SR 185 Rhizothrix minuta DP 159 2954404 Rhizothrix minuta Cletodes limicola DP 264 Rhizothrix minuta SR 249 Rhizothrix minuta SR 163 Rhizothrix minuta SR 164 Rhizothrix minuta SR 165 Rhizothrix minuta SR 250 Rhizothrix minuta DP 100 Rhizothrix minuta DP 160 Haloschizopera pygmaea DP 111 2954408 Haloschizopera pygmaea Haloschizopera pygmaea SR 100 Haloschizopera pygmaea SR 033 Cletodes limicola DP 293 Haloschizopera pygmaea DP 237 Haloschizopera pygmaea DP 166 Haloschizopera pygmaea SR 179 Haloschizopera pygmaea DP 110 Haloschizopera pygmaea SR 402 Haloschizopera pygmaea SR 188 Haloschizopera pygmaea SR 235 Haloschizopera pygmaea DP 109 Haloschizopera pygmaea SR 604 Haloschizopera pygmaea SR 605 Cletodes limicola DP 296 Haloschizopera pygmaea DP 170 Haloschizopera cf pygmaea SR 401 Haloschizopera n sp SR 415 2953922 Haloschizopera Haloschizopera n sp SR 418 Haloschizopera n sp SR 414 Protopsammotopa norvegica SR 201 2955189 Protopsammotopa norvegica Protopsammotopa norvegica SR 246 Protopsammotopa norvegica SR 200 Psammotopa n sp SR 619 2954373 Psammotopa Psammotopa n sp SR 620 Cletodes limicola DP 266 Psammotopa n sp DP 205 Psammotopa n sp DP 206 Eoschizopera n sp aff syltensis SR 413 2954092 Eoschizopera Robergurneya cf spinulosa SR 399 2954696 Robertgurneya spinulosa Bulbamphiascus imus SR 282 2950721 Bulbamphiascus imus Bulbamphiascus imus SR 283 Bulbamphiascus imus DP 238 Bulbamphiascus imus SR 206 Arenopontia cf subterranea SR 580 2954759 Arenopontia subterranea Arenopontia cf subterranea SR 582 Evansula n sp SR 330 2953217 Evansula Arenopontia cf subterranea SR 708 Arenopontia cf subterranea SR 669 Arenopontia cf subterranea DP 302 Arenopontia cf subterranea DP 300 Arenopontia cf subterranea SR 599 Proameira n sp DP 297 2951061 Proameira Proameira n sp DP 313 Proameira n sp DP 314 Proameira n sp DP 315 Proameira n sp SR 647 Evansula cf pygmaea SR 576 2953503 Evansula pygmaea Interleptomesochra cf elongata DP 209 2951099 Leptomesochra elongata Interleptomesochra cf elongata SR 695 Interleptomesochra cf elongata DP 208 Interleptomesochra cf elongata SR 622 Interleptomesochra cf elongata SR 624 Interleptomesochra cf elongata SR 694 Interleptomesochra cf elongata SR 623 Ameira n sp 3 SR 627 1011612 Ameira Sicameira leptoderma DP 210 2950937 Sicameira leptoderma Sicameira leptoderma Copepodidt SR 626 Evansula cf pygmaea SR 578 Sicameira leptoderma SR 342 Sicameira leptoderma SR 203 Sicameira leptoderma SR 204 Filexilia cf intermedia DP 207 2951298 Filexilia intermedia Filexilia cf intermedia SR 621 Filexilia n sp SR 340 2951107 Filexilia Filexilia cf brevipes SR 273 2951297 Filexilia brevipes Filexilia cf brevipes SR 395 Pseudameira crassicornis DP 096 2951306 Pseudameira crassicornis Pseudameira crassicornis DP 097 Evansula cf pygmaea DP 299 Pseudameira crassicornis SR 593 Pseudameira crassicornis SR 594 Pseudameira crassicornis SR 715 Proameira n sp 2 SR 397 2951061 Proameira Proameira n sp 2 SR 400 Proameira n sp 2 SR 396 Proameira n sp 2 SR 398 Proameira n sp 2 SR 405 Ameira n sp SR 696 1011612 Ameira Ameira n sp SR 625 Cletodes longicaudata SR 714 Evansula cf pygmaea SR 654 Ameira n sp SR 628 Ameira n sp DP 211 Ameiridae n sp 2 SR 198 Ameiridae n sp 2 SR 579 Ameira n sp aff parvula 2 SR 409 Ameira n sp aff parvula 2 SR 411 Ameiridae n sp 1 SR 660 Ameira n sp aff parvula 1 SR 221 Ameira n sp aff parvula 1 SR 410 Ameiridae n sp 3 SR 608 Mesochra pygmaea DP 318 2951933 Mesochra pygmaea Ameiridae n sp 3 DP 141 Ameiridae n sp 3 DP 140 Ameiridae n sp 3 SR 609 Ameiridae n sp 3 SR 689 Ameiridae n sp 3 SR 690 Harpacticus flexus DP 006 2951642 Harpacticus flexus Harpacticus flexus SR 176 Harpacticus flexus SR 457 Harpacticus flexus SR 175 Harpacticus flexus SR 454 Mesochra pygmaea SR 709 Harpacticus flexus SR 474 Acrenhydrosoma perplexa SR 707 2949721 Acrenhydrosoma perplexum Enhydrosoma curticauda DP 009 2949682 Enhydrosoma curticauda Enhydrosoma curticauda SR 097 Enhydrosoma curticauda DP 081 Enhydrosoma curticauda SR 015 Enhydrosoma curticauda SR 018 Enhydrosoma curticauda DP 008 Enhydrosoma curticauda DP 087 Enhydrosoma curticauda DP 089 Mesochra pygmaea DP 317 Enhydrosoma curticauda DP 010 Enhydrosoma gariene DP 013 2949757 Enhydrosoma gariene Enhydrosoma gariene SR 484 Enhydrosoma gariene DP 011 Enhydrosoma gariene DP 004 Enhydrosoma gariene SR 479 Enhydrosoma gariene DP 005 Enhydrosoma gariene DP 012 Enhydrosoma gariene SR 480 Enhydrosoma propinquum SR 082 2949713 Enhydrosoma propinquum Mesochra pygmaea DP 316 Enhydrosoma propinquum SR 491 Enhydrosoma propinquum SR 081 Enhydrosoma propinquum SR 489 Enhydrosoma propinquum SR 487 Enhydrosoma propinquum SR 113 Enhydrosoma propinquum SR 486 Enhydrosoma propinquum DP 032 Enhydrosoma propinquum SR 490 Enhydrosoma propinquum SR 539 Enhydrosoma propinquum SR 540 Mesochra pygmaea SR 014 Enhydrosoma propinquum SR 541 Paramesochridae sp 8 SR 663 304676 Paramesochridae Stylicletodes longicaudata DP 286 2949782 Stylicletodes longicaudatus Stylicletodes longicaudatus 2 SR 212 Stylicletodes longicaudatus 2 SR 216 Delavalia palustris DP 017 2955216 Delavalia palustris Delavalia palustris SR 447 Delavalia palustris SR 448 Delavalia palustris SR 473 Delavalia palustris SR 468 Mesochra pygmaea SR 428 Delavalia palustris SR 469 Delavalia palustris SR 471 Delavalia palustris SR 472 Delavalia palustris DP 015 Delavalia palustris DP 016 Delavalia palustris SR 470 Delavalia reflexa SR 088 2955218 Delavalia reflexa Delavalia reflexa SR 138 Beatricella aemula SR 158 2954843 Beatricella aemula Beatricella aemula SR 159 Cylindropsyllus cf remanei SR 285 2954950 Cylindropsyllus remanei Beatricella aemula SR 195 2954843 Beatricella aemula Beatricella aemula SR 194 Beatricella aemula DP 093 Beatricella aemula DP 094 Beatricella aemula SR 157 Amphiascopsis cinctus SR 259 2954393 Amphiascopsis cinctus Amphiascopsis cinctus SR 260 Amphiascopsis cinctus SR 688 Amphiascopsis cinctus DP 135 Amphiascopsis cinctus DP 134 Cylindropsyllus cf remanei SR 306 2954950 Cylindropsyllus remanei Amphiascopsis cinctus DP 139 Ectinosomatidae sp 12 DP 247 296842 Ectinosomatidae Ectinosomatidae sp 12 DP 257 Ectinosomatidae sp 12 DP 256 Ectinosomatidae sp 12 SR 297 Ectinosomatidae sp 12 DP 246 Ectinosomatidae sp 12 DP 255 Ectinosomatidae sp 12 SR 665 Ectinosomatidae sp 12 SR 666 Ectinosomatidae sp 11 DP 194 Cylindropsyllus laevis SR 322 2955041 Cylindropsyllus laevis Ectinosomatidae sp 13 SR 271 Ectinosomatidae sp 31 DP 188 Ectinosomatidae sp 31 SR 686 Ectinosomatidae sp 31 DP 278 Ectinosomatidae sp 31 DP 189 Ectinosomatidae sp 31 DP 187 Ectinosomatidae sp 31 DP 242 Ectinosomatidae sp 24 DP 168 Ectinosomatidae sp 24 DP 184 Ectinosomatidae sp 36 SR 595 Cletodes longicaudata DP 294 Cylindropsyllus laevis SR 321 Arenosetella germanica SR 353 2952500 Arenosetella germanica Arenosetella germanica SR 352 Arenosetella germanica SR 354 Arenosetella germanica SR 355 Ectinosomatidae sp 17 SR 374 Ectinosomatidae sp 27 DP 272 Ectinosomatidae sp 27 SR 648 Ectinosomatidae sp 27 SR 649 Ectinosomatidae sp 27 DP 273 Ectinosomatidae sp 16 DP 182 Cylindropsyllus laevis SR 406 Ectinosomatidae sp 16 DP 183 Ectinosomatidae sp 14 SR 588 Ectinosomatidae sp 14 DP 249 Ectinosomatidae sp 14 SR 296 Ectinosomatidae sp 14 SR 290 Ectinosomatidae sp 14 SR 289 Ectinosomatidae sp 14 SR 295 Ectinosomatidae sp 30 DP 281 Ectinosomatidae sp 5 SR 556 Ectinosomatidae sp 5 SR 535 Leptopontia cf flandrica SR 210 2954261 Leptopontia flandrica Ectinosomatidae sp 5 SR 560 Ectinosomatidae sp 5 SR 512 Ectinosomatidae sp 5 SR 533 Ectinosomatidae sp 5 SR 515 Ectinosomatidae sp 3 SR 509 Ectinosomatidae sp 3 SR 537 Ectinosomatidae sp 3 SR 557 Ectinosomatidae sp 3 SR 527 Ectinosomatidae sp 3 SR 558 Ectinosomatidae sp 3 SR 510 Leptopontia cf flandrica SR 239 Ectinosomatidae sp 2 DP 224 Ectinosomatidae sp 2 SR 511 Ectinosomatidae sp 2 SR 554 Ectinosomatidae sp 2 SR 531 Ectinosomatidae sp 2 SR 530 Ectinosomatidae sp 2 SR 555 Ectinosomatidae sp 2 SR 529 Ectinosomatidae sp 2 SR 559 Ectinosomatidae sp 18 DP 279 Ectinosomatidae sp 18 DP 259 Enhydrosoma sarsi SR 114 2949762 Enhydrosoma sarsi Ectinosomatidae sp 18 DP 239 Ectinosomatidae sp 18 DP 190 Ectinosomatidae sp 18 DP 191 Ectinosomatidae sp 15 DP 130 Ectinosomatidae sp 15 DP 131 Ectinosomatidae sp 15 DP 283 Ectinosomatidae sp 15 SR 267 Ectinosomatidae sp 15 SR 269 Ectinosomatidae sp 15 SR 270 Ectinosomatidae sp 15 SR 268 Enhydrosoma sarsi SR 115 Ectinosomatidae sp 10 DP 221 Ectinosomatidae sp 10 DP 234 Ectinosomatidae sp 10 DP 235 Ectinosomatidae sp 28 DP 280 Ectinosomatidae sp 34 DP 204 Ectinosomatidae sp 35 DP 244 Ectinosomatidae sp 35 SR 662 Ectinosomatidae sp 26 DP 243 Ectinosomatidae sp 26 DP 304 Ectinosomatidae sp 9 SR 567 Enhydrosoma sarsi DP 102 Ectinosomatidae sp 9 SR 661 Ectinosomatidae sp 9 SR 678 Ectinosomatidae sp 23 DP 126 Ectinosomatidae sp 23 DP 127 Ectinosomatidae sp 20 SR 590 Ectinosomatidae sp 20 DP 129 Ectinosomatidae sp 20 DP 186 Ectinosomatidae sp 20 DP 128 Ectinosomatidae sp 20 DP 185 Ectinosomatidae sp 22 SR 601 Arenocaris reducta SR 596 Ectinosomatidae sp 19 SR 367 Ectinosomatidae sp 19 SR 375 Ectinosomatidae sp 19 SR 368 Ectinosomatidae sp 19 SR 366 Ectinosomatidae sp 1 SR 521 Ectinosomatidae sp 1 SR 519 Ectinosomatidae sp 1 SR 552 Ectinosomatidae sp 1 SR 550 Ectinosomatidae sp 1 SR 553 Ectinosomatidae sp 1 SR 551 Arenocaris reducta SR 597 2954550 Arenocaris reducta Ectinosomatidae sp 1 SR 547 Ectinosomatidae sp 1 SR 549 Ectinosomatidae sp 1 SR 546 Ectinosomatidae sp 1 SR 507 Ectinosomatidae sp 1 SR 506 Ectinosomatidae sp 1 SR 548 Ectinosomatidae sp 1 SR 503 Ectinosomatidae sp 1 SR 501 Ectinosomatidae sp 25 DP 201 Ectinosomatidae sp 25 SR 618 Arenocaris reducta SR 218 Ectinosomatidae sp 8 SR 292 Ectinosomatidae sp 8 SR 294 Ectinosomatidae sp 8 DP 231 Ectinosomatidae sp 8 DP 250 Ectinosomatidae sp 8 DP 241 Ectinosomatidae sp 8 SR 291 Ectinosomatidae sp 8 SR 300 Ectinosomatidae sp 8 SR 301 Ectinosomatidae sp 8 SR 307 Ectinosomatidae sp 8 SR 314 Cletodes longicaudata SR 640 Arenocaris reducta DP 326 Ectinosomatidae sp 8 SR 315 Ectinosomatidae sp 8 SR 316 Ectinosomatidae sp 8 DP 222 Ectinosomatidae sp 8 DP 254 Ectinosomatidae sp 8 DP 252 Ectinosomatidae sp 8 DP 192 Ectinosomatidae sp 8 SR 302 Ectinosomatidae sp 8 SR 653 Ectinosomatidae sp 8 DP 193 Ectinosomatidae sp 8 DP 251 Arenocaris reducta DP 325 Ectinosomatidae sp 8 DP 232 Ectinosomatidae sp 8 SR 563 Ectinosomatidae sp 8 DP 228 Ectinosomatidae sp 8 SR 667 Ectinosomatidae sp 8 SR 668 Ectinosomatidae sp 8 DP 229 Ectinosomatidae sp 8 DP 253 Ectinosomatidae sp 8 SR 610 Euterpina acutifrons SR 258 Euterpina acutifrons SR 433 926399 Euterpina acutifrons Arenocaris reducta SR 598 Euterpina acutifrons SR 436 Euterpina acutifrons SR 438 Euterpina acutifrons DP 199 Euterpina acutifrons SR 328 Microarthridion fallax SR 498 2950662 Microarthridion fallax Microarthridion fallax SR 499 Microarthridion fallax SR 496 Altheuta interrupta SR 280 2952025 Alteutha interrupta Danielssenia typica SR 712 2954182 Danielssenia typica Danielsennia typica DP 225 Arenocaris reducta DP 324 Danielssenia typica SR 098 Danielssenia typica SR 162 Danielssenia typica SR 161 Danielssenia typica SR 232 Danielsennia typica DP 282 Danielssenia typica SR 190 Normanella obscura SR 137 2954880 Normanella obscura Normanella obscura SR 635 Normanella obscura SR 035 Normanella obscura SR 145 Arenocaris reducta DP 113 2954550 Arenocaris reducta Normanella obscura SR 012 Normanella mucronata DP 284 2955015 Normanella mucronata Normanella mucronata DP 287 Normanella mucronata SR 633 Normanella mucronata SR 634 Normanella mucronata DP 310 Normanella mucronata DP 311 Arenocaris reducta DP 323 Tachidiella minuta DP 309 2953818 Tachidiella minuta Tachidiella minuta SR 155 Tachidiella minuta DP 307 Tachidiella minuta DP 308 Cletodes longicaudata SR 174 Tachidiella minuta SR 152 Tachidiella minuta SR 154 Tachidiella minuta DP 226 Tachidiella minuta SR 704 Tachidiella minuta SR 153 Tachidiella minuta SR 156 Zosime atlantica SR 637 Zosime atlantica SR 638 Zosime atlantica DP 289 Zosime atlantica DP 288 Cletodes longicaudata DP 295 Zosime atlantica DP 290 Paramesochridae sp 2 SR 631 Paramesochridae sp 2 DP 217 Paramesochridae sp 2 DP 218 Paramesochridae sp 2 DP 215 Paramesochridae sp 2 DP 216 Paramesochridae sp 4 SR 685 Paramesochridae sp 7 SR 583 Paramesochridae sp 6 SR 585 Paramesochridae sp 1 SR 613 Cletodes longicaudata DP 265 Paramesochridae sp 1 SR 615 Paramesochridae sp 1 DP 197 Paramesochridae sp 1 SR 614 Paramesochridae sp 1 SR 611 Paramesochridae sp 1 SR 612 Paramesochridae sp 5 SR 632 Paramesochridae sp 5 SR 700 Metacyclopina harpacticoidea DP 212 Metacyclopina harpacticoidea SR 629 Metacyclopina harpacticoidea DP 214 Cletodes longicaudata SR 217 Metacyclopina harpacticoidea SR 630 Metacyclopina harpacticoidea DP 213 Canuella perplexa SR 034 Canuella perplexa SR 224 Canuella perplexa DP 248 Canuella perplexa SR 187 Canuella perplexa SR 183 Canuella perplexa SR 223 Canuella perplexa DP 240 Canuella perplexa DP 245 ot:bootstrapValues Neighbor-joining node1 ot:bootstrapValues node1 false true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true 0 0 1089 0 1090 546 2 1 2