('Aulacocyclus tambourinensis',((('Passalus lunaris',(('Passalus abortivus',('Paxillus leachi',('Passalus bucki',('Passalus interruptus',('Passalus interstitialis',('Passalus punctiger','Passalus coniferus')))))),(('Passalus jansoni',('Passalus convexus','Passalus ruehli')),(('Passipassalus nukak',('Passalus curtus',('Passalus rugosus','Passalus alfari'))),('Passalus inops',((('Passalus rufiventris','Passalus caelatus'),('Passalus clypeoneleus','Ptichopus angulatus')),('Passalus punctatostriatus',('Ameripassalus guatemalensis',('Passalus cognatus','Passalus spiniger'))))))))),(('Proculus goryi','Proculus mniszechi'),(((('Undulifer acapulcae','Undulifer nigidiodes'),('Pseudacanthus subopacus','Pseudacanthus junctistriatus')),((('Vindex sculptilis','Xylopassaloides chortii'),(('Vindex agnoscendus','Vindex gonzaloi'),'Proculejus sp')),((('Ogyges aluxi',('Ogyges laevissimus','Ogyges crassulus')),('Ogyges championi','Ogyges tzutuhili')),('Oileus bifidus',('Oileus sargi',('Oileus gasparilomi','Oileus rimator')))))),((('Chondrocephalus purulensis',('Chondrocephalus salvadorae',('Chondrocephalus guatemalae',('Chondrocephalus debilis',('Chondrocephalus granulifrons','Chondrocephalus gemmae'))))),((('Arrox agassizi',('Verres corticicola',(('Verres hageni','Verres cavicollis'),('Verres onorei','Verres deficiens')))),('Veturius oberthuri',(('Veturius tuberculifrons','Veturius talamacaensis'),('Veturius aspina',('Veturius crassus','Veturius assimilis'))))),(('Odontotaenius zodiacus',('Spurius bicornis',(('Yumtaax imbellis','Yumtaax recticornis'),('Yumtaax jimenezi','Yumtaax mazatecus')))),(('Popilius gibbosus',('Petrejoides orizabae',('Popilius marginatus','Petrejoides haagi'))),(('Petrejoides tenuis','Petrejoides subrecticornis'),(('Heliscus eclipticus',('Heliscus tropicus','Heliscus vazquezae')),('Pseudoarrox caldasi',('Odontotaenius striatopunctatus','Odontotaenius disjunctus')))))))),('Tonantzin tepetl','Pseudacanthus aztecus'))))),('Leptaulax dentatus',('Erionomus planiceps',('Pharochilus politus','Pharochilus nitidulus')))));