pythonVersion 3.10.12 pythonImplementation CPython true Animals true Source data for tree(s) 'tree1' tree1 ^ot:ottId ^ot:comment highestMintedElementIDs ^ot:nodeLabelMode ^ot:nodeLabelTimeUnit ^ot:reasonsToExcludeFromSynthesis jdduke24 278108 Chondrichthyes ot_3055 Stein, R. W., Mull, C. G., Kuhn, T. S., Aschliman, N. C., Davidson, L. N. K., Joy, J. B., Smith, G. J., Dulvy, N. K., & Mooers, A. O. (2018). Global priorities for conserving the evolutionary history of sharks, rays and chimaeras. Nature Ecology & Evolution, 2(2), 288–298. https://doi.org/10.1038/s41559-017-0448-4 2018 Acroteriobatus annulatus 7060713 Acroteriobatus annulatus Alopias vulpinus 1077204 Alopias vulpinus Carcharhinus leucas 343172 Carcharhinus leucas Carcharhinus limbatus 350158 Carcharhinus limbatus Carcharhinus longimanus 597826 Carcharhinus longimanus Carcharhinus macloti 649215 Carcharhinus macloti Carcharhinus melanopterus 517931 Carcharhinus melanopterus Carcharhinus obscurus 400238 Carcharhinus obscurus Carcharhinus perezii 147228 Carcharhinus perezii Carcharhinus plumbeus 400246 Carcharhinus plumbeus Carcharhinus sealei 805298 Carcharhinus sealei Carcharhinus signatus 902487 Carcharhinus signatus Amblyraja badia 1012668 Amblyraja badia Carcharhinus sorrah 1030721 Carcharhinus sorrah Carcharhinus tilstoni 1030724 Carcharhinus tilstoni Carcharias taurus 331725 Carcharias taurus Carcharodon carcharias 554297 Carcharodon carcharias Centrophorus atromarginatus 805285 Centrophorus atromarginatus Centrophorus granulosus 754130 Centrophorus granulosus Centrophorus harrissoni 104684 Centrophorus harrissoni Centrophorus isodon 805282 Centrophorus isodon Centrophorus moluccensis 449788 Centrophorus moluccensis Centrophorus squamosus 449785 Centrophorus squamosus Amblyraja doellojuradoi 658566 Amblyraja doellojuradoi Centrophorus zeehaani 90311 Centrophorus zeehaani Centroscyllium fabricii 1027221 Centroscyllium fabricii Centroscyllium granulatum 521336 Centroscyllium granulatum Centroscyllium nigrum 1003422 Centroscyllium nigrum Centroscyllium ritteri 1003425 Centroscyllium ritteri Centroscymnus coelolepis 304656 Centroscymnus coelolepis Centroscymnus owstonii 745562 Centroscymnus owstoni Centroselachus crepidater 449809 Centroselachus crepidater Cephaloscyllium albipinnum 805311 Cephaloscyllium albipinnum Cephaloscyllium hiscosellum 90315 Cephaloscyllium hiscosellum Amblyraja hyperborea 619899 Amblyraja hyperborea Cephaloscyllium laticeps 665161 Cephaloscyllium laticeps Cephaloscyllium silasi 769180 Cephaloscyllium silasi Cephaloscyllium umbratile 481015 Cephaloscyllium umbratile Cephaloscyllium variegatum 805306 Cephaloscyllium variegatum Cephaloscyllium ventriosum 224122 Cephaloscyllium ventriosum Cetorhinus maximus 73671 Cetorhinus maximus Chaenogaleus macrostoma 561372 Chaenogaleus macrostoma Chiloscyllium griseum 196309 Chiloscyllium griseum Chiloscyllium hasseltii 178577 Chiloscyllium hasseltii Chiloscyllium indicum 178580 Chiloscyllium indicum Amblyraja jenseni 11779 Amblyraja jenseni Chiloscyllium plagiosum 962561 Chiloscyllium plagiosum Chiloscyllium punctatum 364162 Chiloscyllium punctatum Chimaera fulva 776000 Chimaera fulva Chimaera lignaria 129347 Chimaera lignaria Chimaera monstrosa 29489 Chimaera monstrosa Chimaera opalescens 73641 Chimaera opalescens Chimaera panthera 129350 Chimaera panthera Chimaera phantasma 194652 Chimaera phantasma Chlamydoselachus anguineus 1093624 Chlamydoselachus anguineus Cirrhigaleus asper 68843 Cirrhigaleus asper Amblyraja radiata 513579 Amblyraja radiata Cirrhigaleus australis 24489 Cirrhigaleus australis Cirrhigaleus barbifer 449782 Cirrhigaleus barbifer Cruriraja hulleyi 781245 Cruriraja hulleyi Cruriraja parcomaculata 397100 Cruriraja parcomaculata Dalatias licha 1027232 Dalatias licha Dasyatis akajei 225332 Dasyatis akajei Dasyatis americana 827416 Hypanus americanus Dasyatis bennetti 718969 Hemitrygon bennettii Dasyatis brevicaudata 178571 Dasyatis brevicaudata Dasyatis centroura 648168 Bathytoshia centroura Anoxypristis cuspidata 97265 Anoxypristis cuspidata Dasyatis chrysonota 849622 Dasyatis chrysonota Dasyatis dipterura 960239 Hypanus dipterurus Dasyatis fluviorum 399890 Hemitrygon fluviorum Dasyatis izuensis 61230 Dasyatis izuensis Dasyatis laevigata 61226 Dasyatis laevigata Dasyatis laosensis 815079 Dasyatis laosensis Dasyatis lata 917251 Dasyatis lata Dasyatis longa 100182 Hypanus longus Dasyatis margarita 153861 Fontitrygon margarita Dasyatis margaritella 837115 Fontitrygon margaritella Apristurus albisoma 517911 Apristurus albisoma Dasyatis matsubarai 396786 Dasyatis matsubarai Dasyatis microps 614604 Dasyatis microps Dasyatis navarrae 265174 Dasyatis navarrae Dasyatis parvonigra 129326 Dasyatis parvonigra Dasyatis pastinaca 849605 Dasyatis pastinaca Dasyatis sabina 351766 Dasyatis sabina Dasyatis say 1024125 Dasyatis say Dasyatis thetidis 471820 Dasyatis thetidis Dasyatis ushiei 129331 Dasyatis ushiei Dasyatis zugei 97270 Telatrygon zugei Apristurus ampliceps 364914 Apristurus ampliceps Deania calcea 449777 Deania calcea Deania hystricosa 832506 Deania hystricosa Deania profundorum 769193 Deania profundorum Deania quadrispinosa 3595271 Deania quadrispinosa Dentiraja lemprieri 6139304 Dentiraja lemprieri Diplobatis ommata 773792 Diplobatis ommata Dipturus australis 399883 Dipturus australis Dipturus batis 881959 Dipturus batis Dipturus cerva 399877 Dipturus cerva Dipturus confusus 129357 Dipturus confusus Apristurus aphyodes 303072 Apristurus aphyodes Dipturus gudgeri 129339 Dipturus gudgeri Dipturus healdi 248309 Dipturus healdi Dipturus innominatus 449394 Dipturus innominatus Dipturus kwangtungensis 444672 Dipturus kwangtungensis Dipturus laevis 736066 Dipturus laevis Dipturus leptocaudus 248305 Dipturus leptocauda Dipturus macrocauda 444677 Dipturus macrocauda Dipturus oxyrinchus 884616 Dipturus oxyrinchus Dipturus pullopunctata 397105 Dipturus pullopunctatus Dipturus springeri 264865 Dipturus springeri Acroteriobatus variegatus 7060714 Acroteriobatus variegatus Apristurus australis 281613 Apristurus australis Dipturus tengu 257061 Dipturus tengu Dipturus trachydermus 1014990 Dipturus trachyderma Echinorhinus brucus 521344 Echinorhinus brucus Echinorhinus cookei 1026510 Echinorhinus cookei Etmopterus bigelowi 624105 Etmopterus bigelowi Etmopterus brachyurus 624099 Etmopterus brachyurus Etmopterus dianthus 370409 Etmopterus dianthus Etmopterus dislineatus 426669 Etmopterus dislineatus Etmopterus fusus 426666 Etmopterus fusus Etmopterus gracilispinis 720843 Etmopterus gracilispinis Apristurus brunneus 786098 Apristurus brunneus Etmopterus granulosus 975973 Etmopterus granulosus Etmopterus lucifer 423439 Etmopterus lucifer Etmopterus molleri 624144 Etmopterus molleri Etmopterus polli 426663 Etmopterus polli Etmopterus princeps 832502 Etmopterus princeps Etmopterus pseudosqualiolus 524155 Etmopterus pseudosqualiolus Etmopterus pusillus 651004 Etmopterus pusillus Etmopterus schultzi 370395 Etmopterus schultzi Etmopterus sentosus 426656 Etmopterus sentosus Etmopterus sheikoi 624137 Etmopterus sheikoi Apristurus exsanguis 456193 Apristurus exsanguis Etmopterus spinax 1059232 Etmopterus spinax Etmopterus splendidus 105820 Etmopterus splendidus Etmopterus unicolor 592868 Etmopterus unicolor Etmopterus virens 426653 Etmopterus virens Eucrossorhinus dasypogon 274889 Eucrossorhinus dasypogon Euprotomicrus bispinatus 547470 Euprotomicrus bispinatus Eusphyra blochii 1021212 Eusphyra blochii Figaro boardmani 105487 Figaro boardmani Furgaleus macki 665114 Furgaleus macki Galeocerdo cuvier 976355 Galeocerdo cuvier Apristurus fedorovi 517909 Apristurus fedorovi Galeorhinus galeus 898984 Galeorhinus galeus Galeus arae 654149 Galeus arae Galeus atlanticus 742155 Galeus atlanticus Galeus eastmani 296640 Galeus eastmani Galeus melastomus 327712 Galeus melastomus Galeus murinus 647388 Galeus murinus Galeus polli 264875 Galeus polli Galeus sauteri 821054 Galeus sauteri Ginglymostoma cirratum 400227 Ginglymostoma cirratum Glaucostegus cemiculus 745043 Glaucostegus cemiculus Apristurus japonicus 1095199 Apristurus japonicus Glaucostegus thouin 1924 Glaucostegus thouin Glaucostegus typus 463561 Glaucostegus typus Glyphis fowlerae 1024581 Glyphis fowlerae Glyphis gangeticus 1024585 Glyphis gangeticus Glyphis garricki 19949 Glyphis garricki Glyphis glyphis 741692 Glyphis glyphis Gollum attenuatus 1037442 Gollum attenuatus Gymnura altavela 225323 Gymnura altavela Gymnura australis 836401 Gymnura australis Gymnura crebripunctata 244283 Gymnura crebripunctata Apristurus kampae 1095202 Apristurus kampae Gymnura japonica 759548 Gymnura japonica Gymnura marmorata 674426 Gymnura marmorata Gymnura micrura 244274 Gymnura micrura Gymnura poecilura 651021 Gymnura poecilura Gymnura zonura 618803 Aetoplatea zonura Halaelurus lineatus 839454 Halaelurus lineatus Halaelurus natalensis 17080 Halaelurus natalensis Halaelurus sellus 1024578 Halaelurus sellus Haploblepharus edwardsii 201988 Haploblepharus edwardsii Harriotta raleighana 776006 Harriotta raleighana Apristurus laurussonii 303074 Apristurus laurussonii Hemigaleus australiensis 651018 Hemigaleus australiensis Hemigaleus microstoma 90191 Hemigaleus microstoma Hemipristis elongata 19955 Hemipristis elongata Hemiscyllium ocellatum 516305 Hemiscyllium ocellatum Hemitriakis complicofasciata 800582 Hemitriakis complicofasciata Hemitriakis falcata 651015 Hemitriakis falcata Hemitriakis japanica 1037427 Hemitriakis japanica Hemitriakis leucoperiptera 1024575 Hemitriakis leucoperiptera Heptranchias perlo 281365 Heptranchias perlo Heterodontus francisci 1019121 Heterodontus francisci Apristurus longicephalus 1095189 Apristurus longicephalus Heterodontus galeatus 651027 Heterodontus galeatus Heterodontus mexicanus 228673 Heterodontus mexicanus Heterodontus portusjacksoni 1019118 Heterodontus portusjacksoni Heterodontus zebra 654133 Heterodontus zebra Hexanchus griseus 435193 Hexanchus griseus Hexatrygon bickelli 103243 Hexatrygon bickelli Himantura astra 142175 Himantura astra Himantura fai 399899 Pateobatis fai Himantura gerrardi 674420 Himantura gerrardi Himantura granulata 323635 Urogymnus granulatus Apristurus macrorhynchus 1037436 Apristurus macrorhynchus Himantura hortlei 180434 Pateobatis hortlei Himantura imbricata 815070 Himantura imbricata Himantura jenkinsii 836406 Pateobatis jenkinsii Himantura leoparda 357806 Himantura leoparda Himantura lobistoma 781238 Himantura lobistoma Himantura oxyrhyncha 1024570 Himantura oxyrhyncha Himantura pacifica 151438 Himantura pacifica Himantura pastinacoides 180437 Maculabatis pastinacoides Himantura polylepis 815047 Himantura chaophraya Himantura schmardae 151435 Himantura schmardae Apristurus manis 415535 Apristurus manis Himantura signifer 1022121 Himantura signifer Himantura toshi 836392 Himantura toshi Himantura uarnacoides 836395 Pateobatis uarnacoides Himantura uarnak 394365 Himantura uarnak Himantura undulata 648161 Himantura undulata Himantura walga 180431 Himantura walga Holohalaelurus regani 694589 Holohalaelurus regani Hongeo koreana 3595458 Hongeo koreana Hydrolagus affinis 3595471 Hydrolagus affinis Hydrolagus africanus 694594 Hydrolagus africanus Aculeola nigra 550082 Aculeola nigra Apristurus melanoasper 1095192 Apristurus melanoasper Hydrolagus barbouri 886196 Hydrolagus barbouri Hydrolagus bemisi 142160 Hydrolagus bemisi Hydrolagus colliei 29493 Hydrolagus colliei Hydrolagus lemures 261757 Hydrolagus lemures Hydrolagus melanophasma 338748 Hydrolagus melanophasma Hydrolagus mirabilis 3595466 Hydrolagus mirabilis Hydrolagus mitsukurii 394375 Hydrolagus mitsukurii Hydrolagus novaezealandiae 261770 Hydrolagus novaezealandiae Hydrolagus ogilbyi 261761 Hydrolagus ogilbyi Hydrolagus trolli 180443 Hydrolagus trolli Apristurus microps 303081 Apristurus microps Hypogaleus hyugaensis 748408 Hypogaleus hyugaensis Iago garricki 498761 Iago garricki Iago omanensis 1037433 Iago omanensis Insentiraja subtilispinosa 808372 Insentiraja subtilispinosa Irolita waitii 180440 Irolita waitii Isistius brasiliensis 277019 Isistius brasiliensis Isurus oxyrinchus 73672 Isurus oxyrinchus Isurus paucus 500598 Isurus paucus Lamiopsis temminckii 19944 Lamiopsis temminckii Lamna ditropis 950993 Lamna ditropis Apristurus pinguis 281593 Apristurus pinguis Lamna nasus 500593 Lamna nasus Leucoraja circularis 649762 Leucoraja circularis Leucoraja erinacea 493418 Leucoraja erinacea Leucoraja fullonica 660328 Leucoraja fullonica Leucoraja garmani 781232 Leucoraja garmani Leucoraja melitensis 138103 Leucoraja melitensis Leucoraja naevus 660331 Leucoraja naevus Leucoraja ocellata 734611 Leucoraja ocellata Leucoraja wallacei 192602 Leucoraja wallacei Loxodon macrorhinus 966255 Loxodon macrorhinus Apristurus platyrhynchus 281598 Apristurus platyrhynchus Malacoraja kreffti 431890 Malacoraja kreffti Manta birostris 920351 Mobula birostris Megachasma pelagios 73662 Megachasma pelagios Mitsukurina owstoni 73664 Mitsukurina owstoni Mobula hypostoma 476719 Mobula hypostoma Mobula japanica 180418 Mobula japanica Mobula kuhlii 71599 Mobula kuhlii Mobula munkiana 498773 Mobula munkiana Mobula tarapacana 180423 Mobula tarapacana Mobula thurstoni 674430 Mobula thurstoni Apristurus profundorum 1027218 Apristurus profundorum Mustelus antarcticus 976346 Mustelus antarcticus Mustelus asterias 208184 Mustelus asterias Mustelus californicus 47455 Mustelus californicus Mustelus canis 976343 Mustelus canis Mustelus henlei 1035182 Mustelus henlei Mustelus lenticulatus 1026206 Mustelus lenticulatus Mustelus lunulatus 841889 Mustelus lunulatus Mustelus manazo 573813 Mustelus manazo Mustelus mosis 1035178 Mustelus mosis Mustelus mustelus 208178 Mustelus mustelus Apristurus sinensis 90303 Apristurus sinensis Mustelus norrisi 902535 Mustelus norrisi Mustelus palumbes 171731 Mustelus palumbes Mustelus punctulatus 208181 Mustelus punctulatus Mustelus ravidus 498769 Mustelus ravidus Mustelus schmitti 1035185 Mustelus schmitti Mustelus sinusmexicanus 192852 Mustelus sinusmexicanus Mustelus stevensi 583753 Mustelus stevensi Mustelus widodoi 215760 Mustelus widodoi Myliobatis aquila 114274 Myliobatis aquila Myliobatis australis 261767 Myliobatis australis Aptychotrema rostrata 233377 Aptychotrema rostrata Myliobatis californicus 47684 Myliobatis californica Myliobatis freminvillei 781226 Myliobatis freminvillei Myliobatis goodei 1018989 Myliobatis goodei Myliobatis longirostris 100179 Myliobatis longirostris Myliobatis tobijei 298364 Myliobatis tobijei Narcine brevilabiata 295468 Narcine brevilabiata Narcine brunnea 385698 Narcine brunnea Narcine entemedor 808382 Narcine entemedor Narcine lasti 703724 Narcine lasti Narcine tasmaniensis 73023 Narcine tasmaniensis Aptychotrema vincentiana 281618 Aptychotrema vincentiana Narcine timlei 920356 Narcine timlei Narcine westraliensis 703721 Narcine westraliensis Nasolamia velox 720835 Nasolamia velox Nebrius ferrugineus 833437 Nebrius ferrugineus Negaprion acutidens 517928 Negaprion acutidens Negaprion brevirostris 450137 Negaprion brevirostris Neoharriotta pinnata 195190 Neoharriotta pinnata Neoraja caerulea 745026 Neoraja caerulea Neotrygon annotata 592850 Neotrygon annotata Neotrygon kuhlii 688899 Neotrygon kuhlii Arhynchobatis asperrimus 771294 Arhynchobatis asperrimus Neotrygon leylandi 399902 Neotrygon leylandi Neotrygon ningalooensis 323624 Neotrygon ningalooensis Neotrygon picta 545646 Neotrygon picta Notoraja azurea 808388 Notoraja azurea Notorynchus cepedianus 290409 Notorynchus cepedianus Odontaspis noronhai 756794 Odontaspis noronhai Okamejei acutispina 745029 Okamejei acutispina Okamejei boesemani 1081601 Okamejei boesemani Okamejei cairae 813666 Okamejei cairae Okamejei hollandi 808378 Okamejei hollandi Asymbolus parvus 805296 Asymbolus parvus Okamejei jensenae 353506 Okamejei jensenae Okamejei kenojei 167107 Okamejei kenojei Okamejei meerdervoortii 725199 Okamejei meerdervoortii Orectolobus floridus 798336 Orectolobus floridus Orectolobus hutchinsi 703731 Orectolobus hutchinsi Orectolobus japonicus 798346 Orectolobus japonicus Orectolobus maculatus 572746 Orectolobus maculatus Orectolobus ornatus 516310 Orectolobus ornatus Orectolobus parvimaculatus 798343 Orectolobus parvimaculatus Oxynotus bruniensis 817093 Oxynotus bruniensis Aetobatus flagellum 767872 Aetobatus flagellum Asymbolus rubiginosus 805294 Asymbolus rubiginosus Oxynotus centrina 99815 Oxynotus centrina Oxynotus paradoxus 956140 Oxynotus paradoxus Paragaleus pectoralis 813401 Paragaleus pectoralis Parascyllium collare 154119 Parascyllium collare Paratrygon aiereba 674438 Paratrygon ajereba Parmaturus xaniurus 568234 Parmaturus xaniurus Pastinachus atrus 818497 Pastinachus atrus Pastinachus gracilicaudus 703734 Pastinachus gracilicaudus Pastinachus sephen 815094 Pastinachus sephen Pastinachus solocirostris 178574 Pastinachus solocirostris Atelomycterus baliensis 541147 Atelomycterus baliensis Pastinachus stellurostris 813324 Pastinachus stellurostris Pavoraja alleni 801138 Pavoraja alleni Pavoraja nitida 1931 Pavoraja nitida Platyrhina sinensis 920361 Platyrhina sinensis Platyrhinoidis triseriata 519270 Platyrhinoidis triseriata Plesiobatis daviesi 444671 Plesiobatis daviesi Plesiotrygon iwamae 151414 Plesiotrygon iwamae Pliotrema warreni 12069 Pliotrema warreni Poroderma africanum 553095 Poroderma africanum Poroderma pantherinum 1037439 Poroderma pantherinum Atelomycterus fasciatus 541153 Atelomycterus fasciatus Potamotrygon castexi Potamotrygon falkneri 127560 Potamotrygon falkneri Potamotrygon henlei 127575 Potamotrygon henlei Potamotrygon hystrix 450831 Potamotrygon hystrix Potamotrygon motoro 151418 Potamotrygon motoro Potamotrygon orbignyi 1094314 Potamotrygon orbignyi Potamotrygon yepezi 309617 Potamotrygon yepezi Prionace glauca 361523 Prionace glauca Pristiophorus cirratus 1095624 Pristiophorus cirratus Pristiophorus japonicus 234341 Pristiophorus japonicus Atelomycterus marmoratus 541150 Atelomycterus marmoratus Pristiophorus nudipinnis 754113 Pristiophorus nudipinnis Pristis clavata 703742 Pristis clavata Pristis pectinata 1025533 Pristis pectinata Pristis pristis 3594619 Pristis pristis Pristis zijsron 703739 Pristis zijsron Proscyllium habereri 821048 Proscyllium habereri Proscymnodon plunketi 139131 Proscymnodon plunketi Psammobatis lentiginosa 878126 Psammobatis lentiginosa Psammobatis rudis 618609 Psammobatis rudis Pseudocarcharias kamoharai 647448 Pseudocarcharias kamoharai Atelomycterus marnkalha 541140 Atelomycterus marnkalha Pseudoginglymostoma brevicaudatum 24097 Pseudoginglymostoma brevicaudatum Pseudotriakis microdon 456205 Pseudotriakis microdon Pteromylaeus bovinus 126580 Aetomylaeus bovinus Pteroplatytrygon violacea 478977 Pteroplatytrygon violacea Raja asterias 884658 Raja asterias Raja brachyura 607396 Raja brachyura Raja clavata 493424 Raja clavata Raja inornata 519276 Raja inornata Raja maderensis 692727 Raja maderensis Raja microocellata 607393 Raja microocellata Atlantoraja castelnaui 258171 Atlantoraja castelnaui Raja miraletus 884661 Raja miraletus Raja montagui 1093440 Raja montagui Raja radula 607390 Raja radula Raja rhina 374348 Raja rhina Raja stellulata 519279 Raja stellulata Raja straeleni 840630 Raja straeleni Raja undulata 884629 Raja undulata Raja velezi 126586 Raja velezi Rajella barnardi 397087 Rajella barnardi Rajella bathyphila 995421 Rajella bathyphila Atlantoraja cyclophora 781240 Atlantoraja cyclophora Rajella bigelowi 882967 Rajella bigelowi Rajella caudaspinosa 995438 Rajella caudaspinosa Rajella fyllae 745033 Rajella fyllae Rajella kukujevi 745038 Rajella kukujevi Rajella leopardus 995394 Rajella leopardus Rhina ancylostoma 444665 Rhina ancylostoma Rhincodon typus 214509 Rhincodon typus Rhinobatos annandalei 131247 Rhinobatos annandalei Rhinobatos formosensis 444666 Glaucostegus formosensis Rhinobatos glaucostigma 11768 Pseudobatos glaucostigma Atlantoraja platana 712041 Atlantoraja platana Rhinobatos horkelii 751553 Pseudobatos horkelii Rhinobatos hynnicephalus 520113 Rhinobatos hynnicephalus Rhinobatos jimbaranensis 225288 Rhinobatos jimbaranensis Rhinobatos lentiginosus 550653 Pseudobatos lentiginosus Rhinobatos penggali 695926 Rhinobatos penggali Rhinobatos percellens 751549 Pseudobatos percellens Rhinobatos productus 1095489 Pseudobatos productus Rhinobatos punctifer 769191 Rhinobatos punctifer Rhinobatos rhinobatos 813059 Rhinobatos rhinobatos Rhinobatos schlegelii 970832 Rhinobatos schlegelii Aulohalaelurus labiosus 541144 Aulohalaelurus labiosus Rhinochimaera africana 970830 Rhinochimaera africana Rhinochimaera pacifica 194644 Rhinochimaera pacifica Rhinoptera bonasus 197732 Rhinoptera bonasus Rhinoptera javanica 970840 Rhinoptera javanica Rhinoptera jayakari 801133 Rhinoptera jayakari Rhinoptera neglecta 836389 Rhinoptera neglecta Rhinoptera steindachneri 151425 Rhinoptera steindachneri Rhinoraja albomaculata 84108 Bathyraja albomaculata Rhinoraja macloviana 84127 Bathyraja macloviana Rhinoraja magellanica 605824 Bathyraja magellanica Bathyraja abyssicola 147169 Bathyraja abyssicola Rhinoraja multispinis 605829 Bathyraja multispinis Rhinoraja murrayi 967216 Bathyraja murrayi Rhinoraja taranetzi 914980 Bathyraja taranetzi Rhizoprionodon acutus 272025 Rhizoprionodon acutus Rhizoprionodon lalandii 1083224 Rhizoprionodon lalandii Rhizoprionodon longurio 804022 Rhizoprionodon longurio Rhizoprionodon oligolinx 804019 Rhizoprionodon oligolinx Rhizoprionodon porosus 197740 Rhizoprionodon porosus Rhizoprionodon taylori 1060348 Rhizoprionodon taylori Rhizoprionodon terraenovae 611836 Rhizoprionodon terraenovae Aetobatus narinari 759551 Aetobatus narinari Bathyraja aleutica 147182 Bathyraja aleutica Rhynchobatus australiae 399871 Rhynchobatus australiae Rhynchobatus djiddensis 970835 Rhynchobatus djiddensis Rhynchobatus laevis 984897 Rhynchobatus laevis Rhynchobatus palpebratus 227145 Rhynchobatus palpebratus Rioraja agassizii 588194 Rioraja agassizii Rostroraja alba 257036 Rostroraja alba Schroederichthys bivius 618627 Schroederichthys bivius Scoliodon laticaudus 380998 Scoliodon laticaudus Scoliodon macrorhynchos 458523 Scoliodon macrorhynchos Scyliorhinus canicula 1026500 Scyliorhinus canicula Bathyraja brachyurops 84116 Bathyraja brachyurops Scyliorhinus capensis 806125 Scyliorhinus capensis Scyliorhinus retifer 150264 Scyliorhinus retifer Scyliorhinus stellaris 901864 Scyliorhinus stellaris Scyliorhinus torazame 266108 Scyliorhinus torazame Scylliogaleus quecketti 1035188 Scylliogaleus quecketti Scymnodon ringens 956136 Scymnodon ringens Sinobatis borneensis 618822 Sinobatis borneensis Somniosus antarcticus 1022134 Somniosus antarcticus Somniosus microcephalus 442057 Somniosus microcephalus Somniosus pacificus 547460 Somniosus pacificus Bathyraja cousseauae 84114 Bathyraja cousseauae Somniosus rostratus 821060 Somniosus rostratus Sphyrna corona 577843 Sphyrna corona Sphyrna lewini 450134 Sphyrna lewini Sphyrna media 577849 Sphyrna media Sphyrna mokarran 970846 Sphyrna mokarran Sphyrna tiburo 218666 Sphyrna tiburo Sphyrna tudes 1058817 Sphyrna tudes Sphyrna zygaena 970843 Sphyrna zygaena Squaliolus aliae 956130 Squaliolus aliae Squaliolus laticaudus 69179 Squaliolus laticaudus Bathyraja griseocauda 84132 Bathyraja griseocauda Squalus acanthias 1019126 Squalus acanthias Squalus brevirostris 620710 Squalus brevirostris Squalus chloroculus 211583 Squalus chloroculus Squalus crassispinus 1026219 Squalus crassispinus Squalus cubensis 194647 Squalus cubensis Squalus edmundsi 1026216 Squalus edmundsi Squalus grahami 449806 Squalus grahami Squalus hemipinnis 211581 Squalus hemipinnis Squalus japonicus 936746 Squalus japonicus Squalus megalops 350352 Squalus megalops Bathyraja irrasa 967205 Bathyraja irrasa Squalus mitsukurii 194649 Squalus mitsukurii Squalus montalbani 620711 Squalus montalbani Squalus nasutus 1026210 Squalus nasutus Squalus suckleyi 1019106 Squalus suckleyi Squatina aculeata 1028968 Squatina aculeata Squatina africana 1028971 Squatina africana Squatina albipunctata 1028952 Squatina albipunctata Squatina armata 1028957 Squatina armata Squatina australis 399868 Squatina australis Squatina californica 1095501 Squatina californica Bathyraja kincaidii 755991 Bathyraja kincaidii Squatina dumeril 902505 Squatina dumeril Squatina formosa 25734 Squatina formosa Squatina guggenheim 1028961 Squatina guggenheim Squatina japonica 1028964 Squatina japonica Squatina legnota 1028976 Squatina legnota Squatina nebulosa 909554 Squatina nebulosa Squatina occulta 1028981 Squatina occulta Squatina pseudocellata 605493 Squatina pseudocellata Squatina squatina 821063 Squatina squatina Squatina tergocellata 399865 Squatina tergocellata Bathyraja lindbergi 147201 Bathyraja lindbergi Squatina tergocellatoides 605489 Squatina tergocellatoides Stegostoma fasciatum 122336 Stegostoma fasciatum Sutorectus tentaculatus 309097 Sutorectus tentaculatus Sympterygia acuta 1040562 Sympterygia acuta Sympterygia bonapartii 97081 Sympterygia bonapartii Taeniura lymma 151429 Taeniura lymma Taeniurops grabata 1941 Taeniura grabata Taeniurops meyeni 688878 Taeniura meyeni Torpedo californica 493412 Tetronarce californica Torpedo fuscomaculata 232016 Torpedo fuscomaculata Bathyraja maccaini 967202 Bathyraja maccaini Torpedo mackayana 372007 Torpedo mackayana Torpedo macneilli 25737 Tetronarce macneilli Torpedo marmorata 493406 Torpedo marmorata Torpedo nobiliana 346065 Tetronarce nobiliana Torpedo sinuspersici 254411 Torpedo sinuspersici Torpedo tokionis 970849 Tetronarce tokionis Torpedo torpedo 355116 Torpedo torpedo Triaenodon obesus 309093 Triaenodon obesus Triakis megalopterus 748396 Triakis megalopterus Triakis scyllium 32033 Triakis scyllium Bathyraja maculata 389364 Bathyraja maculata Triakis semifasciata 32029 Triakis semifasciata Trigonognathus kabeyai 956126 Trigonognathus kabeyai Trygonoptera imitata 25722 Trygonoptera imitata Trygonoptera mucosa 25740 Trygonoptera mucosa Trygonoptera ovalis 25743 Trygonoptera ovalis Trygonoptera personata 25719 Trygonoptera personata Trygonoptera testacea 178594 Trygonoptera testacea Trygonorrhina fasciata 463555 Trygonorrhina fasciata Typhlonarke aysoni 802673 Typhlonarke aysoni Urobatis concentricus 819000 Urobatis concentricus Bathyraja mariposa 389367 Bathyraja mariposa Urobatis halleri 18608 Urolophus halleri Urobatis jamaicensis 450826 Urobatis jamaicensis Urobatis maculatus 227128 Urolophus maculatus Urogymnus asperrimus 970804 Urogymnus asperrimus Urolophus aurantiacus 970801 Urolophus aurantiacus Urolophus bucculentus 178588 Urolophus bucculentus Urolophus cruciatus 836383 Urolophus cruciatus Urolophus expansus 178591 Urolophus expansus Urolophus flavomosaicus 178566 Urolophus flavomosaicus Urolophus kapalensis 178569 Urolophus kapalensis Aetomylaeus maculatus 836386 Aetomylaeus maculatus Bathyraja meridionalis 967219 Bathyraja meridionalis Urolophus lobatus 546882 Urolophus lobatus Urolophus paucimaculatus 836398 Urolophus paucimaculatus Urolophus sufflavus 701857 Urolophus sufflavus Urolophus viridis 701854 Urolophus viridis Urolophus westraliensis 836380 Urolophus westraliensis Urotrygon rogersi 569133 Urotrygon rogersi Zameus squamulosus 975969 Zameus squamulosus Zanobatus schoenleinii 126589 Zanobatus schoenleinii Zapteryx brevirostris 228228 Zapteryx brevirostris Zapteryx exasperata 181823 Zapteryx exasperata Bathyraja minispinosa 529898 Bathyraja minispinosa Zearaja nasuta 449397 Dipturus nasutus Bathyraja pallida 771310 Bathyraja pallida Bathyraja parmifera 529910 Bathyraja parmifera Bathyraja richardsoni 967198 Bathyraja richardsoni Bathyraja scaphiops 605834 Bathyraja scaphiops Bathyraja shuntovi 967195 Bathyraja shuntovi Bathyraja smirnovi 824086 Bathyraja smirnovi Bathyraja spinicauda 580254 Bathyraja spinicauda Bathyraja trachura 914977 Bathyraja trachura Aetomylaeus vespertilio 281606 Aetomylaeus vespertilio Bathyraja violacea 824089 Bathyraja violacea Beringraja binoculata 389361 Raja binoculata Beringraja pulchra 7060749 Beringraja pulchra Brachaelurus colcloughi 771316 Brachaelurus colcloughi Brachaelurus waddi 274894 Brachaelurus waddi Brochiraja albilabiata 771319 Brochiraja albilabiata Brochiraja asperula 298412 Brochiraja asperula Brochiraja leviveneta 378168 Brochiraja leviveneta Brochiraja microspinifera 977388 Brochiraja microspinifera Brochiraja spinifera 977358 Brochiraja spinifera Alopias pelagicus 801808 Alopias pelagicus Bythaelurus canescens 803567 Bythaelurus canescens Bythaelurus dawsoni 378153 Bythaelurus dawsoni Callorhinchus callorynchus 550636 Callorhinchus callorynchus Callorhinchus capensis 776015 Callorhinchus capensis Callorhinchus milii 550662 Callorhinchus milii Carcharhinus acronotus 147232 Carcharhinus acronotus Carcharhinus albimarginatus 805301 Carcharhinus albimarginatus Carcharhinus altimus 147235 Carcharhinus altimus Carcharhinus amblyrhynchoides 649208 Carcharhinus amblyrhynchoides Carcharhinus amblyrhynchos 517925 Carcharhinus amblyrhynchos Alopias superciliosus 73666 Alopias superciliosus Carcharhinus borneensis 378195 Carcharhinus borneensis Carcharhinus brachyurus 378279 Carcharhinus brachyurus Carcharhinus brevipinna 517922 Carcharhinus brevipinna Carcharhinus cautus 649205 Carcharhinus cautus Carcharhinus dussumieri 665131 Carcharhinus dussumieri Carcharhinus falciformis 787198 Carcharhinus falciformis Carcharhinus fitzroyensis 649218 Carcharhinus fitzroyensis Carcharhinus galapagensis 378287 Carcharhinus galapagensis Carcharhinus isodon 147225 Carcharhinus isodon Carcharhinus leiodon 829416 Carcharhinus leiodon ot:time Myr Maximum likelihood node1 node1 false true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true 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