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^ot:nodeLabelMode ^ot:nodeLabelTimeUnit rootedge tree2 maxmaronna L-Leclere 1010196 Leptothecata false ot_714 Maximiliano M. Maronna, Thaís P. Miranda, Álvaro L. Peña Cantero, Marcos S. Barbeitos, Antonio C. Marques, 2016, 'Towards a phylogenetic classification of Leptothecata (Cnidaria, Hydrozoa)', Scientific Reports, vol. 6, p. 18075 2016 Solmundella bitentaculata 962896 Solmundella bitentaculata Orthopyxis everta 144203 Orthopyxis everta Campanularia hincksii 144201 Campanularia hincksii Orthopyxis crenata 479998 Orthopyxis crenata Bonneviella sp4 296263 Bonneviella Bonneviella sp2 296263 Bonneviella Bonneviella sp3 296263 Bonneviella Bonneviella regia 296264 Bonneviella regia Campanularia volubilis 296262 Campanularia volubilis Rhizocaulus verticillatus 296260 Rhizocaulus verticillatus Campanularia sp 4712825 Campanularia Stauridiosarsia ophiogaster 141204 Stauridiosarsia ophiogaster Silicularia rosea 144208 Silicularia rosea Gonothyraea loveni 446122 Gonothyraea loveni Laomedea calceolifera 446121 Laomedea calceolifera Obelia geniculata 508706 Obelia geniculata Obelia bidentata 782818 Obelia bidentata Obelia longissima 854448 Obelia longissima Obelia dichotoma 1097118 Obelia dichotoma Laomedea flexuosa 259585 Laomedea flexuosa Gonothyraea inornata 782820 Laomedea inornata Clytia hummelincki 554369 Clytia hummelincki Sarsia lovenii 896966 Sarsia lovenii Clytia paulensis 554371 Clytia paulensis Clytia noliformis 714002 Clytia noliformis Clytia hemisphaerica 33386 Clytia hemisphaerica Clytia hemisphaerica2 33386 Clytia hemisphaerica Clytia linearis 554370 Clytia linearis Clytia gracilis 554368 Clytia gracilis Eirene brevistylus 1085438 Eirene brevistylus Staurotheca pachyclada 709319 Staurotheca pachyclada Staurotheca nonscripta 709317 Staurotheca nonscripta Staurotheca compressa 709329 Staurotheca compressa Coryne pintneri 3681284 Coryne pintneri Staurotheca antarctica 709327 Staurotheca antarctica Abietinella operculata 287313 Abietinella operculata Zygophylax biarmata 829370 Zygophylax biarmata Cryptolaria pectinata 392866 Cryptolaria pectinata Antarctoscyphus grandis 135113 Antarctoscyphus grandis Antarctoscyphus elongatus 800697 Antarctoscyphus elongatus Antarctoscyphus spiralis 709324 Antarctoscyphus spiralis Symplectoscyphus curvatus 185719 Symplectoscyphus curvatus Symplectoscyphus liouvillei 304210 Symplectoscyphus liouvillei Symplectoscyphus exochus 760035 Symplectoscyphus exochus Pennaria disticha 707213 Pennaria disticha Symplectoscyphus plectilis 304223 Symplectoscyphus plectilis Symplectoscyphus glacialis 304258 Symplectoscyphus glacialis Symplectoscyphus tricuspidatus 631138 Symplectoscyphus tricuspidatus Halecium halecinum 1027946 Halecium halecinum Halecium pusillum 1096597 Halecium pusillum Halecium beanii 1027947 Halecium beanii Halecium lankesterii 1027945 Halecium lankesteri Halecium mirabile 3682421 Halecium mirabile Halecium petrosum 889800 Halecium petrosum Halecium incertus 185730 Halecium incertus Cladocoryne floccosa 132725 Cladocoryne floccosa Halecium delicatulum 185732 Halecium delicatulum Halecium muricatum 735255 Halecium muricatum Halecium mediterraneum 1096599 Halecium mediterraneum Halecium lenticulare 1096601 Halecium lenticulare Halecium labrosum 735256 Halecium labrosum Halecium tenellum 502043 Halecium tenellum Sertularella mediterranea 971191 Sertularella mediterranea Sertularella ellisii 971201 Sertularella ellisii Sertularella gayi 971189 Sertularella gayi Sertularella sanmatiasensis 185721 Sertularella sanmatiasensis Porpita porpita 288598 Porpita porpita Scandia gigas 109534 Scandia gigas Sertularella diaphana 4713274 Sertularella diaphana Sertularella robusta 3683487 Sertularella robusta Sertularella polyzonias 502051 Sertularella polyzonias Symplectoscyphus turgidus 971195 Symplectoscyphus turgidus Sertularella tenella 3683399 Sertularella tenella Sertularella africana 971199 Sertularella africana Sertularella rugosa 631132 Sertularella rugosa Thyroscyphus ramosus 3683567 Thyroscyphus ramosus Thyroscyphus marginatus 480016 Thyroscyphus marginatus Zanclea prolifera 427245 Zanclea prolifera Sertularelloides cylindritheca 271834 Sertularella cylindritheca Sertularia turbinata 3683110 Sertularia turbinata Sertularia tumida 3683154 Sertularia tumida Sertularia marginata 196815 Sertularia marginata Sertularia perpusilla 315300 Sertularia perpusilla Sertularia unguiculata 58259 Sertularia unguiculata Idiellana pristis 3683687 Idiellana pristis Dynamena moluccana 444958 Dynamena moluccana Dynamena disticha 631137 Dynamena disticha Dynamena obliqua 3683192 Dynamena obliqua Moerisia inkermanica 982739 Moerisia inkermanica Dynamena crisioides 3683189 Dynamena crisioides Dynamena pumila 58258 Dynamena pumila Salacia desmoides 444954 Salacia desmoides Diphasia rosacea 1027952 Diphasia rosacea Diphasia fallax 58256 Diphasia fallax Amphisbetia minima 58260 Amphisbetia minima Amphisbetia operculata 570023 Amphisbetia operculata Sertularia distans 271832 Sertularia distans Sertularia cupressina 631134 Sertularia cupressina Sertularia argentea 971193 Sertularia argentea Eudendrium glomeratum 385357 Eudendrium glomeratum Thuiaria cornigera 3683331 Thuiaria cornigera Thuiaria thuja 631136 Thuiaria thuja Abietinaria filicula 58252 Abietinaria filicula Abietinaria abietina 788001 Abietinaria abietina Hydrallmania falcata 58254 Hydrallmania falcata Hydrodendron gardineri 683683 Hydrodendron gardineri Hydrodendron mirabile 769996 Hydrodendron mirabile Nemalecium lighti 3682533 Nemalecium lighti Gymnangium hians 710450 Gymnangium hians Gymnangium montagui 1027948 Gymnangium montagui Maeotias marginata 350088 Maeotias marginata Eudendrium caraiuru 3681717 Eudendrium caraiuru Cladocarpus integer 570027 Cladocarpus integer Lytocarpia canepa 3682619 Lytocarpia canepa Gymnangium gracilicaule 769997 Gymnangium gracilicaule Macrorhynchia philippina 770007 Macrorhynchia philippina Macrorhynchia phoenicea 719384 Macrorhynchia phoenicea Macrorhynchia phoenicea2 719384 Macrorhynchia phoenicea Lytocarpia phyteuma 683681 Lytocarpia phyteuma Lytocarpia sp 683680 Lytocarpia Aglaophenia lophocarpa 392870 Aglaophenia lophocarpa Aglaophenia elongata 570019 Aglaophenia elongata Eudendrium carneum 122693 Eudendrium carneum Aglaophenia acacia 570017 Aglaophenia acacia Aglaophenia parvula 145046 Aglaophenia parvula Aglaophenia octodonta 145045 Aglaophenia octodonta Aglaophenia pluma 145053 Aglaophenia pluma Aglaophenia tubiformis 6501 Aglaophenia tubiformis Aglaophenia harpago 570021 Aglaophenia harpago Aglaophenia picardi 889791 Aglaophenia picardi Aglaophenia kirchenpaueri 515920 Aglaophenia kirchenpaueri Aglaophenia struthionides 3682539 Aglaophenia struthionides Aglaophenia tubulifera 392871 Aglaophenia tubulifera Candelabrum cocksii 183504 Candelabrum cocksii Aglaophenia rhynchocarpa 3682564 Aglaophenia rhynchocarpa Aglaophenia latecarinata 770006 Aglaophenia latecarinata Schizotricha nana 760025 Schizotricha nana Schizotricha trinematotheca 135100 Schizotricha trinematotheca Schizotricha falcata 760023 Schizotricha falcata Schizotricha turqueti 135098 Schizotricha turqueti Schizotricha crassa 760041 Schizotricha crassa Ventromma halecioides 315306 Kirchenpaueria halecioides Ventromma halecioides2 315306 Kirchenpaueria halecioides Pycnotheca mirabilis 719382 Pycnotheca mirabilis Hydra circumcincta 149372 Hydra circumcincta Kirchenpaueria pinnata 6504 Kirchenpaueria pinnata Kirchenpaueria pinnata2 6504 Kirchenpaueria pinnata Oswaldella stepanjantsae 135103 Oswaldella stepanjantsae Oswaldella incognita 800683 Oswaldella incognita Oswaldella grandis 135096 Oswaldella grandis Oswaldella shetlandica 3682833 Oswaldella shetlandica Oswaldella laertesi 135105 Oswaldella laertesi Polyplumaria flabellata 502049 Polyplumaria flabellata Halopteris minuta 6503 Halopteris minuta Halopteris schucherti 167462 Halopteris schucherti Ectopleura larynx 183506 Ectopleura larynx Halopteris tenella 196051 Halopteris tenella Halopteris diaphana 196047 Halopteris diaphana Halopteris alternata 196050 Halopteris alternata Halopteris liechtensternii 566399 Halopteris liechtensternii Halopteris carinata 985076 Halopteris carinata Halopteris catharina 769992 Halopteris catharina Monostaechas quadridens 719394 Monostaechas quadridens Antennella secundaria 770005 Antennella secundaria Antennella ansini 889788 Antennella ansini Antennella kiwiana 145052 Antennella kiwiana Ectopleura obypa 3681781 Ectopleura obypa Dentitheca bidentata 145047 Dentitheca bidentata Plumularia spiralis 683679 Plumularia spiralis Plumularia habereri 145054 Plumularia habereri Plumularia habereri2 145054 Plumularia habereri Plumularia insignis 3682198 Plumularia insignis Plumularia pulchella 329831 Plumularia pulchella Plumularia margaretta 889792 Plumularia margaretta Plumularia margaretta2 889792 Plumularia margaretta Plumularia obliqua 952423 Plumularia obliqua Plumularia filicaulis 145055 Plumularia filicaulis Corymorpha pendula 863655 Corymorpha pendula Plumularia hyalina 6502 Plumularia hyalina Plumularia lagenifera 221333 Plumularia lagenifera Plumularia setacea 566402 Plumularia setacea Nemertesia ventriculiformis 502048 Nemertesia ventriculiformis Nemertesia antennina 631146 Nemertesia antennina Proboscidactyla ornata 805156 Proboscidactyla ornata Fabienna sphaerica 168731 Fabienna sphaerica Hydrichthella epigorgia 231178 Hydrichthella epigorgia Olindias sambaquiensis 717858 Olindias sambaquiensis Hydractinia sp 35792 Hydractinia Adelopora crassilabrum 805153 Adelopora crassilabrum Lizzia blondina 902447 Lizzia blondina Leuckartiara octona 836779 Leuckartiara octona Perarella schneideri 902445 Perarella schneideri Turritopsis nutricula 461175 Turritopsis nutricula Bougainvillia muscus 45501 Bougainvillia muscus Bimeria vestita 975494 Bimeria vestita Physalia physalis 702294 Physalia physalis Apolemia sp 536355 Apolemia Geryonia proboscidalis 386743 Geryonia proboscidalis Praya dubia 486578 Praya dubia Stephalia dilata 863143 Stephalia dilata Cordagalma ordinata 5675976 Cordagalma ordinata Physophora hydrostatica 408506 Physophora hydrostatica Forskalia edwardsii 5675969 Forskalia edwardsii Hincksella sp 4713270 Hincksella Acryptolaria conferta 392868 Acryptolaria conferta Lafoea dumosa 109524 Lafoea dumosa Stegella lobata 598539 Stegella lobata Melicertum octocostatum 696558 Melicertum octocostatum Liriope tetraphylla 1097123 Liriope tetraphylla Billardia subrufa 732085 Billardia subrufa Modeeria rotunda 394804 Modeeria rotunda Stegopoma plicatile 709322 Stegopoma plicatile Laodicea undulata 394796 Laodicea undulata Melicertissa sp 934508 Melicertissa Staurodiscus gotoi 515131 Staurodiscus gotoi Hebella scandens 3682875 Hebella scandens Hebella venusta 1021487 Hebella venusta Anthohebella parasitica 109518 Anthohebella parasitica Mitrocomella brownei 1096593 Mitrocomella brownei Crossota rufobrunnea 883621 Crossota rufobrunnea Mitrocomella niwai 1096591 Mitrocomella niwai Calycella syringa 446143 Calycella syringa Opercularella lacerata 971197 Opercularella lacerata Campanulina panicula 570025 Campanulina panicula Campanulina pumila 598547 Opercularella pumila Phialella quadrata 394801 Phialella quadrata Tiaropsidium kelseyi 54550 Tiaropsidium kelseyi Tiaropsis multicirrata 1021490 Tiaropsis multicirrata Rhacostoma atlanticum 425701 Rhacostoma atlanticum Eutonina indicans 585723 Eutonina indicans Aglantha digitale 459324 Aglantha digitale Aequorea conica 311788 Aequorea conica Aequorea victoria 1084491 Aequorea victoria Aequorea forskalea 258884 Aequorea aequorea Aequorea coerulescens 1018788 Aequorea coerulescens Octophialucium indicum 315307 Octophialucium indicum Eirene viridula 570029 Eirene viridula Aequorea floridana 1038041 Aequorea floridana Blackfordia virginica 766639 Blackfordia virginica Sugiura chengshanense 442 Sugiura chengshanense Eirene pyramidalis 812832 Eirene pyramidalis Pantachogon haeckeli 985083 Pantachogon haeckeli Eirene hexanemalis 812841 Eirene hexanemalis Eirene lacteoides 812845 Eirene lacteoides Eirene kambara 812839 Eirene kambara Helgicirrha malayensis 288889 Helgicirrha malayensis Helgicirrha brevistyla 288887 Helgicirrha brevistyla Hydranthea margarica 196048 Hydranthea margarica Eucheilota menoni 1096607 Eucheilota menoni Eucheilota maculata 1096609 Eucheilota maculata Eutima curva 1096605 Eutima curva Eutima gegenbauri 1096603 Eutima gegenbauri Botrynema brucei 827332 Botrynema brucei Eutima krampi 812830 Eutima krampi Eutima levuka 288891 Eutima levuka Eutima sapinhoa 425766 Eutima sapinhoa Eugymnanthea japonica 229364 Eugymnanthea japonica Eirene ceylonensis 812835 Eirene ceylonensis Eirene menoni 812843 Eirene menoni Lovenella gracilis 446098 Lovenella gracilis Eucheilota bakeri 446110 Eucheilota bakeri Orthopyxis sargassicola 144196 Orthopyxis sargassicola Orthopyxis integra 144192 Orthopyxis integra ot:substitutionCount Maximum likelihood node89 ot:posteriorSupport tree2_ROOT false true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true false true true true false true true true true true true true true true true true true true true true true true true true false true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true false true true true true true true true true true true true true true true true true true true true true true true true true false true true true true false true true true true true true true true true true true true true true true true true true true true true true true false true true true true true false true true true true true true true true 1 1 0.99 1 0.49 0.98 0.91 1 0.88 1 0.5 0.87 0.91 0.61 0.07 0.48 0.35 0.8 1 0.2 0.83 1 0.78 0.99 1 1 0.32 0.23 0.19 0.13 0.31 0.12 0.26 0.87 0.12 1 0.56 0.45 0.27 0.39 0.29 0.98 0.54 0.93 0.89 0.8 0.98 0.23 0.58 0.87 0.95 0.25 0.1 0.83 0.2 1 0.72 0.59 1 0.18 0.38 0.94 0.61 0.36 1 0.49 0.72 0.59 0.22 0.13 0.05 0.07 0.18 0.27 0.22 0.87 0.57 0.37 0.13 0.87 0.96 0.59 0.3 0.35 0.89 0.99 0.86 0.5 0.58 0.97 0.99 0.83 1 0.61 0.98 0.29 0.99 0.38 0.19 0.26 0.9 0.46 0.65 0.34 0.32 0.31 0.52 0.64 0.58 0.59 0.8 1 0.91 0.93 0.02 0.91 0.48 0.96 0.7 0.87 0.95 0.83 0.65 0.33 0.18 0.97 0.59 1 0.69 0.55 0.25 0.99 0.51 0.9 0.97 0.86 0.11 0.37 0.34 0.44 0.76 1 0.45 0.63 0.78 0.97 0.85 0.19 0.28 0.14 0.71 0.99 0.24 0.41 0.1 0.55 0.67 0.87 0.06 0.08 0.17 0.55 0.38 0.94 0.04 0.28 0.92 0.76 0.67 0.86 0.4 1 0.11 0.27 0.4 0.03 0.2 0.05 0.13 0.11 0.07 0.08 0.81 0.41 1 0.31 0.89 0.28 0.62 0.94 0.84 0.47 0.45 0.19 0.58 0.99 1 0.62 0.69 0.63 0.79 0.86 0.95 0.41 0.81 1 0.99 0.06 0.95 0.07 0.46 0.82 0.5 0.08 0.12 0.31 0.46 0.43 0.79 1 0.63 0.46 0.44 0.7 0.78 1 1 0.82 0.46 1 1 0.39 0.02 0.14 0.99 0.44 1 0.01 1 0.63 1 0.99 0.76 0.54 1 0.62 1 0.83 1 0.49 0.55 0.55 1 0.99 1 0.57 0.62 0.7 0.65 0.86 ot:substitutionCount Maximum likelihood node614 ot:bootstrapValues node528 false true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true 97 100 100 100 100 100 35 100 97 100 78 89 76 68 81 32 19 51 100 9 47 100 75 100 100 100 21 100 46 50 60 66 37 13 18 70 48 68 37 59 100 99 25 94 41 44 98 36 80 99 99 41 4 100 10 100 8 35 87 76 78 100 95 100 44 48 43 63 80 83 31 49 10 24 25 16 9 15 95 13 93 94 73 28 79 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true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true true 98 100 100 100 100 100 32 100 95 100 91 83 71 72 83 39 27 12 51 100 57 100 62 100 100 100 24 100 33 34 46 66 41 11 12 58 44 72 37 54 100 97 17 92 41 41 98 82 99 100 54 10 100 25 100 39 75 75 64 100 94 100 66 63 40 59 71 73 23 51 21 10 8 9 19 98 82 47 23 95 95 78 33 84 97 100 96 76 69 100 100 82 100 93 6 15 23 40 44 94 27 57 14 61 54 93 27 100 31 9 92 100 59 10 97 96 98 84 71 87 100 94 90 6 21 99 79 100 99 35 64 100 56 74 22 74 15 91 100 34 43 74 100 99 36 37 45 84 99 1 77 64 18 43 13 12 57 65 100 64 87 20 42 44 100 44 99 79 77 43 99 18 14 15 47 57 20 11 14 98 68 100 29 71 100 91 99 97 97 94 91 43 47 38 100 100 57 37 92 97 98 100 47 82 100 100 100 32 35 97 82 95 67 37 22 23 45 88 100 48 35 5 95 55 100 100 88 19 100 100 16 45 26 99 71 0 7 100 100 61 100 80 80 100 76 100 94 100 77 14 93 100 97 100 94 96 87 96 99